Site function and the ‘ibex-site phenomenon’: myth or reality?

Summary.  This paper focuses on ibex hunting in Italy during the Upper Palaeolithic. The faunal assemblages from five ibex-dominated sites (Dalmeri, Arene Candide, Fumane, Villabruna and Soman) are examined with the aim of assessing the role of the sites and people's activities in the wider settlement system. These sites will be placed in their wider context of the Italian Upper Palaeolithic so as to explore the nature of variability in ibex exploitation. The importance of these considerations is to examine aspects of specialized hunting, increased intensity of resource exploitation during the end of the Upper Palaeolithic, and methodological considerations of the relationship between faunal remains and the human activities that produced them.     

Epipaleolithic and Mesolithic adaptations in Cantabrian Spain and Pyrenean France

The period of deglaciation from ca. 13,000 to ca. 9000 B.P. along the northern edge of the Cantabrian Cordillera and Pyrenees was characterized by marked climatic and environmental oscillations, culminating in the establishment of interglacial conditions. While along the Cantabrian coast, late Upper Paleolithic groups had long been developing diversified systems of adaptation, fully exploiting the wide range of food resources of that narrow but ecologically varied region (notably red deer and marine mollusks), Magdalenian hunters along the southern edge of the Aquitaine basin were becoming increasingly specialized in the hunting of one medium-size game species, reindeer. Thus, while the artifact industries and artistic traditions of the two adjacent regions along the forty-third parallel developed along similar lines in the Magdalenian and Azilian, and despite a common montane specialization in ibex hunting, the changes that came with the end of the Last Glacial affected the human groups of the two regions very differently, as reflected in the early Mesolithic records of Vasco-Cantabria and Gascony, respectively.     

Rabbits as Food at the end of the Upper Palaeolithic at Molí del Salt (Catalonia,Spain)

Many factors have been causally linked to the diversification of hunting during the European Palaeolithic: declining supplies of high‐ranked prey, considerable human demographic growth, reduced residential mobility, larger populations of ubiquitous small mammals and significant technological developments. However, small prey exploitation was not uniform: the European rabbit (Oryctolagus cuniculus ) is the most frequent species in the Upper Palaeolithic archaeological record of the Iberian Peninsula – south and Mediterranean area – and Southern France. This is demonstrated at Molí del Salt, an Upper Palaeolithic site located at Vimbodí (Catalonia, Spain), whose mammal fauna stands out for the predominance of rabbits [91% of minimum number of individuals (n = 136)]. We analysed the faunal remains from one level [A^sup (c. 12 700–13 000 cal BP)] in order to identify the agent responsible for the faunal accumulation, and to reconstruct aspects of procurement and consumption that shed light on Palaeolithic subsistence strategies in the Northeast Iberian Peninsula. Our results indicate that human agency rather than carnivore activity was responsible for the bone accumulation at Molí del Salt. We identified all the stages in the consumption sequence from skinning to ingestion. We argue that the rabbits were mostly harvested during summer or winter or both seasons. Clearly, the European rabbit was a target species for the human groups which lived at Molí del Salt providing meat, and skin. Copyright © 2016 John Wiley & Sons, Ltd.     

Reindeer Hunting in the Upper Palaeolithic: Sex Ratios as a Reflection of Different Procurement Strategies

This paper is a first attempt to investigate the variability in reindeer-procurement strategies in Upper Palaeolithic sites by means of sex ratios, as reflected by osteometrical data. For this purpose the “Variability Size Index” method is employed. The idea behind the analysis is that sex ratios in reindeer (but also in red deer and in bovids) were mainly determined—as they are in modern populations—by the reproductive biology of the animals and not by environmental conditions. Thus, any deviations from these ratios must be explained in terms of behavioural/cultural variables. The faunal material for the study comes from sites in southwest France, Switzerland and southern Germany. During the Upper Palaeolithic females dominate in all assemblages, but a clear variability can be seen: some sites show sex ratios similar to those found in Palaeolithic carnivore dens, whereas other sites show somewhat higher frequencies of bulls. The variation in the sex ratios of reindeer among Upper Palaeolithic sites cannot be explained by either their chronological position or topographical location, but it is more probably a result from seasonal differences in reindeer-procurement. Thus generalizations about the exploitation of reindeer or other prey species based on results of a single site or layer of a site will mask the variability in the subsistence strategies which existed either during the Upper Palaeolithic or, at a finer scale, even during the Magdalenian.     

Hard core and cutting edge: experimental manufacture and use of Magdalenian composite projectile tips

The technology of the European Upper Palaeolithic yielded abundant evidence of the use of composite projectile heads, in the form of osseous points on the side of which one or several (micro)lithic elements are attached. Yet, little experimental work has been devoted to testing and assessing the parameters of use of this type of composite tips. In this paper we present a pilot experiment with replicas of Magdalenian composite spear tips, made of an antler point with one or two rows of flint backed bladelets. Two series of replicas were manufactured after the lithic and osseous record of, respectively, the Lower Magdalenian from southwest France (c. 20–18 Ky cal BP) and the Upper Magdalenian of Pincevent in the Paris Basin (c. 15–14 Ky cal BP). The 34 experimental composite heads were hafted to spears that were then shot with a spearthrower at the carcasses of two young deer. The results provide some insight into the performance characteristics of the osseous and lithic components, both in efficiency and durability. Finally, possible improvements of the experimental protocol are discussed, as well as the implications of our results for the understanding of projectile point variability in the Upper Palaeolithic.     

The Upper Paleolithic Rock Art of Iberia

Nearly 200 rock art sites of Upper Paleolithic age are currently known on the Iberian Peninsula, in both caves and the open air. Over half are still concentrated in Cantabrian Spain and they span the period between c. 30–11 kya, but–tracking the course of human demography in this geographically circumscribed region–many of the images were probably painted or engraved during the Solutrean and, especially, Magdalenian. Dramatic discoveries and dating projects have significantly expanded the Iberian rock art record both geographically and temporally in recent years, in close coincidence with the growth of contemporaneous archeological evidence: cave art loci in Aragón and Levante attributable to the Solutrean and Magdalenian, many cave art sites and a few open-air ones in Andalucía and Extremadura that are mostly Solutrean (in line with evidence of a major Last Glacial Maximum human refugium in southern Spain), the spectacular Côa Valley open-air complex in northern Portugal (together with a growing number of other such loci and one cave) that was probably created during the Gravettian-Magdalenian periods, and a modest, but important increase in proven cave and open-air sites in the high, north-central interior of Spain that are probably Solutrean and/or Magdalenian. Despite regional variations in decorated surfaces, themes, techniques and styles, there are broad (and sometimes very specific) pan-Iberian similarities (as well as ones with the Upper Paleolithic art of southern France) that are indicative of widespread human contacts and shared systems of symbols and beliefs during the late Last Glacial. As this Ice Age world and the forms of social relationships and ideologies that helped human groups survive in it came to an end, so too did the decoration of caves, rockshelters and outcrops, although in some regions other styles of rock art would return under very different conditions of human existence.     

The role of shellfish in hunter–gatherer societies during the Early Upper Palaeolithic: A view from El Cuco rockshelter,northern Spain

The role of coastal resources in the subsistence strategies of Palaeolithic human populations has only recently become an important topic in Old World archaeology. Information on the exploitation of these resources, both as foodstuffs and symbolic elements, can be used to infer the emergence of modern human behaviour as well as to track the diversification and intensification of human diet over time. The excavations carried out at El Cuco rockshelter, located in northern Spain have provided evidence for the exploitation of marine resources during the Early Upper Palaeolithic. The accumulation of Patella shells at El Cuco provides the largest accumulation and the first clear evidence of collection and consumption of molluscs during the Aurignacian on the Atlantic Façade of Europe. A deposit of ornamental shells appeared in a very homogeneous context dated to the Gravettian, suggesting that the shells belonged to the same item. The analysis of this evidence has allowed us to conclude that marine resources were systematically used at least from the beginning of the Upper Palaeolithic in the region. However, a comparison with the Mesolithic shows that intensive shellfish gathering did not occur until the end of the Upper Palaeolithic. Regarding the ornaments, it is interpreted that the identified shell beads were used as social or personal status markers.     

Seasonality and age structure in an archaeological assemblage of Sika deer (Cervus nippon)

Sika deer (Cervus nippon) was one of the major terrestrial mammals hunted in Japan throughout the Jomon period, which extends from the end of the Palaeolithic until the arrival of rice agriculture in the first millennium bc . Mandibular analysis of hunted deer is believed to be more useful in determining the season of death than antlers in archaeological contexts. This paper aims to present a new method of estimating an occupational season of a Jomon site by using sika deer mandibles. This method consists of two stages. First, two measurements on a mandible should be taken. If the distribution shows any gaps among young juveniles, this strongly suggests that the sika was hunted in a specific season. Second, tooth eruption and attrition stage should be observed; this may suggest the season in which hunting took place. This method was utilized to analyse sika deer remains from Awashimadai in eastern Kanto. The result of the analysis shows that deer were hunted only from spring to summer at Awashimadai. Putting these results together with the estimate of seasonality deduced from other species strongly suggests that the site was occupied only from April to July. This paper also examines how the tooth attrition stages of sika deer progress. The preliminary comparison with red deer (Cervus elaphus) in England suggests that molar wear progresses more quickly in sika than in red deer. Copyright © 1999 John Wiley & Sons, Ltd.     

On stone-boiling technology in the Upper Paleolithic: behavioral implications from an Early Magdalenian hearth in El Mirón Cave,Cantabria, Spain

Stone boiling is one of the principal cooking methods used by hunter-gatherer societies. The present paper proposes behavioral and organizational inferences as to how stone boiling was incorporated into hunter-gatherer subsistence practices through an examination of a shallow-basin hearth in an Early Magdalenian level (c. 15,500 ¹⁴C B.P.) of El Mirón Cave, Cantabria (northern Spain). Exploratory analysis of spatial patterns of archaeological remains (bones, lithic artifacts, and fire-cracked rocks) and use-life analysis of fire-cracked rocks demonstrate that the hearth was used and maintained during visits of humans who preyed mainly on ibex and red deer near the site. The relative accessibility of these ungulates and cost-induced technology of stone boiling suggest the implication that stone boiling was employed to maximize the energy and nutrition obtained from carcasses of these game taxa under the circumstance of resource intensification.     

Evidence from the Yana Palaeolithic site,Arctic Siberia,yields clues to the riddle of mammoth hunting

It has become commonplace to talk about humans hunting mammoths, and overhunting is thought to have been one of the causes of the mammoth extinction. However, definite evidence of mammoth kills by humans remains surprisingly scarce. Here we show convincing evidence of mammoth hunting in the Siberian Arctic between 29 000 and 27 000 ¹⁴C years BP. Our data set, from the Yana Upper Palaeolithic site (Siberian Arctic), includes the following: fragments of lithic points and ivory shaft embedded in two mammoth scapulae; two identical holes made by projectiles in a mammoth scapula and a pelvic bone; mammoth tongue bones found in the cultural layer far away from the main mammoth bone accumulation, indicating the consumption of fresh mammoth meat; and a narrow mammoth bone size distribution, implying hunting selection based on animal size. The data suggest that Palaeolithic Yana humans hunted mammoths sporadically, presumably when ivory was needed for making tools. Such non-intensive hunting practiced by humans over millennia would not be fatal to a sustainable mammoth population.     

Taphonomic and Methodological Perspectives of Leporid Hunting During the Upper Paleolithic of the Western Mediterranean Basin

Leporid (rabbit and hare) bones have been shown to yield important information about subsistence practices, mobility patterns, and demographic trends during the Paleolithic of the western and eastern Mediterranean regions. Studies of Spanish Paleolithic caves rich in rabbit bones suggest that residential mobility patterns influence the degree of leporid hunting through time. Studies of Paleolithic sites in the eastern Mediterranean suggest that leporids were hunted in large numbers only after population sizes and densities reached certain thresholds. This paper reviews and critiques these studies based on current taphonomic and ecologic information about leporids. Leporid hunting during the Upper Paleolithic of central Portugal is then discussed and compared to these existing models. These latter data suggest that rabbit hunting in central Portugal does not conform to any existing model, suggesting that local factors of leporid density and environmental conditions likely influenced the nature and timing of small game acquisition during the Upper Paleolithic.     

Mammalian fauna from Palaeolithic sites in the Upper Yenisei River basin (southern Siberia): Review of the current zooarchaeological evidence

The palaeontological data on mammal remains from two Middle Palaeolithic (ca. 125 000 years ago – 27 200 BP) and 34 (ca. 34 000–12 000 BP) Upper Palaeolithic cultural complexes in the upper part of the Yenisei River basin (southern Siberia) are considered. General features of the faunal assemblages are established. Several issues are discussed, including (a) changes in species composition through time and palaeoenvironmental implications of the zooarchaeological records; (b) patterns of human exploitation (hunting) of mammals; (c) issue of mammoth hunting and (d) possibility of domestication of dog in the late Upper Palaeolithic in the Yenisei River basin. Copyright © 2009 John Wiley & Sons, Ltd.     

MAKING A DIFFERENCE: PALAEOLITHIC ICONOGRAPHY AS A TRAIT OF IDENTITY IN THE IBERIAN PENINSULA

The study of the figurative repertoire of Palaeolithic artists allows us to approach aspects such as iconographic diffusion and cultural preferences. This paper presents an updated corpus of figurative rock art for the Iberian peninsula and analyses its distribution in the Cantabrian region, inland Iberia and the Mediterranean basin, three areas frequently used in the literature. This corpus contains 3341 graphic units that were analysed using multivariate statistics and hypothesis testing. The results show that the main figurative themes can be classified according to their discriminating power. Horse, ibex and deer were the animal motifs that formed the common background of this artistic expression. In contrast, Palaeolithic artists used different proportions of bison, hind and aurochs to create a possible distinctive identity. Finally, it is discussed whether the iconographic selection is due to palaeoecological reasons, cultural motivations, or a combination of both.     

Probable evidences of bone accumulation by Pleistocene bearded vulture at the archaeological site of El Mirón Cave (Spain)

The bearded vulture (Gypaetus barbatus), a bird or prey that mainly feeds on bones of large mammal carcasses, has been recently evidenced as an accumulating agent with significant contributions to palaeontological assemblages in caves and shelters, but extremely rare at archaeological sites. The results of the taphonomic analyses carried out in the Upper Palaeolithic layers of El Mirón Cave (Cantabrian Spain), which are presented here, evidence the existence of special digestive marks and a typical skeletal pattern in a noticeable amount of small and medium-sized ungulates bones. This fact suggest that a representative part of the bone assemblage has been caused by bearded vultures, and strengthens the idea that this bird has to be definitely included among other potential non-human accumulators in archaeological sites.     

Magdalenian Children: Projectile Points,Portable Art and Playthings

Children, no doubt, were a significant component of Upper Palaeolithic societies. Despite this fact, however, serious identification and consideration of material culture which may have belonged to children – at least at one time during their use‐life – have not been undertaken. This situation extends to the best represented and most intensively studied of the European Palaeolithic techno‐complexes, the Magdalenian (c.21,000–14,000 cal BP), and consequently, we know very little about the children of this enigmatic people. As play, including object play, is a ‘true cultural universal’, we can be certain that Magdalenian children integrated objects into their games, with these playthings later incorporated into the archaeological record. Through examining ethnographic accounts of recent hunter‐gatherer children and reconsidering archaeological assemblages in light of these data, this paper suggests that Magdalenian playthings probably included full‐sized adult weapon tips and – more significantly – pieces of what archaeologists term ‘art mobilier’.     

Sexual dimorphism in limb bones of ibex (Capra ibex L.): mixture analysis applied to modern and fossil data

Estimating sex ratios of fossil bone assemblages is an important step in the determination of demographic profiles, which are essential for understanding the palaeobiology and palaeoethology of any particular species, as well as its exploitation patterns by humans. This is especially true for ibex (Capra ibex), which was a main source of food for hominids during Pleistocene times. Classical methods for determining sexual dimorphism and sex ratio, such as analyses using uni‐ and bivariate plots, are based on an arbitrary fixing of limits between sexes. Here we use a more robust statistical method termed mixture analysis (MA) to determine the sex of postcranial remains (long bones, metapodials and tarsals) from ibex. For the first time, we apply MA to both a modern and a fossil sample of one species, by using metric data taken from (i) a collection of present‐day ibex skeletons and (ii) a Palaeolithic sample of the same species. Our results clearly show that the forelimb (humerus and radius) is more dimorphic than the hindlimb (femur and tibia) and is therefore better suited for sexing ibex. It also appears that metapodials should be used carefully for estimating sex ratios. On the basis of these results, we propose a classification of bone measurements that are more or less reliable for sexing ibex. The results of MA applied to the ibex fossil bones from the Upper Palaeolithic site of the Observatoire (Monaco) lead us to the conclusion that this assemblage consists of a majority of males. The quantitative estimations calculated by the MA make it possible to compare the size of Pleistocene and modern ibex for the whole set of variables used in this study. Copyright © 2006 John Wiley & Sons, Ltd.     

Specialized Early Upper Palaeolithic Hunters in Southwestern France?

Paul Mellars has long used cave and rockshelter ungulate faunal assemblages from southwestern France to argue that the early Upper Palaeolithic people of this region focused their hunting on reindeer (Rangifer tarandus), and that such specialized hunting distinguishes the Upper from the Middle Palaeolithic in at least this region. We examine this argument quantitatively, using a sample of 133 Mousterian, Châtelperronian, and Aurignacian ungulate assemblages. We show that only five Aurignacian assemblages, from three sites, stand out in terms of the degree to which their ungulate faunas are dominated by a single taxon. We also show that some Mousterian cave and rock shelter ungulate assemblages are more heavily dominated by large bovids than Aurignacian assemblages are dominated by reindeer, and that Mellars' argument is highly dependent on the exclusion of open sites from the analysis and on the numerical threshold he has selected to indicate hunting specialization.     

Taphonomy and zooarchaeology of a high-altitude Upper Pleistocene faunal sequence from Hovk-1 Cave,Armenia

The Caucasus is a key region for the study of hominid evolution and Neanderthal ecology. Taphonomic and zooarchaeological studies of sites from this region are few and only focused on sites at low-to-mid altitude zones with evidence of relatively intensive hominid occupation. This study focused on the taphonomic and zooarchaeological characteristics of a high-altitude site from the Upper Pleistocene – Hovk-1 Cave – looking at diachronic change in both natural and cultural processes which shaped the faunal assemblage. Results best fit a model in which the bones of most large mammals, mainly ungulates (wild goat, Capra aegagrus and red deer, Cervus elaphus) and cave bears (Ursus spelaeus) accumulated naturally through pitfalls, with minimal input from human or carnivore activity. This accumulation is characterized by a high frequency of complete ungulate and carnivore bones, a bear assemblage which is dominated by young-adults and a wild goat assemblage that includes juvenile and young-adult individuals. Our taphonomic reconstruction serves as a point of reference for comparative studies of palaeoenvironments and human subsistence patterns of Middle and Upper Palaeolithic sites in the Caucasus and broadens our perspective on hominid occupation and ecological adaptation in other high-altitude world regions.     

Taphonomy and zooarchaeology of the Upper Palaeolithic cave of Dzudzuana,Republic of Georgia

We present the results of a detailed taphonomic and zooarchaeological study of the faunal remains from the Upper Palaeolithic layers of Dzudzuana Cave, Republic of Georgia. This study presents the first carefully analysed Upper Palaeolithic faunal assemblage from the southern Caucasus and thus serves as a significant point of reference for inter‐regional studies of Upper Palaeolithic subsistence in Eurasia. A series of intra‐site taphonomic comparisons are employed to reconstruct the depositional history of the bone assemblages within the different occupational phases at the site and to investigate subsistence, meat procurement and bone‐processing strategies. Caucasian tur (Capra caucasica), aurochs (Bos primigenius) and steppe bison (Bison priscus) were the major prey species throughout the Upper Palaeolithic. Their frequencies do not change significantly over time, and nor does bone preservation vary by layer. The assemblage is characterised by significant density‐mediated biases, caused by both human bone‐processing behaviours and in situ post‐burial bone attrition. Bone marrow extraction produced large numbers of unidentified bone fragments, many exhibiting green bone fractures. The density and size of bone assemblages and the extent of fragmentation indicate that Dzudzuana Cave was repeatedly occupied by Upper Palaeolithic foragers over many years. Skeletal part representation and butchery marks from all stages of carcass processing suggest that prey occasionally underwent field butchery. Intra‐site taphonomic comparisons highlight uniform patterns of cultural and economic behaviours related to food procurement and processing strategies. Copyright © 2007 John Wiley & Sons, Ltd.