Insect invaders of reconstructed Anglo-Saxon houses at West Stow,Suffolk, England

Abstract

The insect fauna, particularly beetles (Coleoptera), of four modern reconstructions of Anglo-Saxon houses at West Stow, Suffolk, UK, has been investigated by pitfall trapping. Modest numbers of insects were recovered from each structure. The overlap with the fauna recovered from archaeological floor deposits of about the same period was notable, being greater than predicted, since the reconstructions lacked the litter and waste believed to have been available as insect habitats in the past.     

Species Associations Among Insect Remains from Urban Archaeological Deposits and their Significance in Reconstructing the Past Human Environment

Assemblages of archaeological insect (mainly Coleoptera) remains from in and around Anglo-Scandinavian buildings at 16–22 Coppergate, York, were investigated in order to identify associations between pairs and groups of species, and to determine the significance of those associations as indicators of past human activity and living conditions. Following initial exploration, a pairwise measure of association was used to establish working groups of co-occurring species, which were then related to their likely habitats. Detrended canonical correspondence analysis (DCCA) was used to test the groups. Robust and consistent species associations were identified among the death assemblages. The groups defined by the analyses usually corresponded to the ecological preferences of species, although some groups of species would not be found together at the present day. Some groups probably represented complexes of habitats that occurred together in the past, and not single communities, and some may be peculiar to Coppergate and similar sites. It is suggested that the species associations are of value in assemblage interpretation, and that variations in associations among sites will reflect archaeologically significant aspects of the sites.     

Pitfalls in the environmental interpretation of insect death assemblages

Attention is drawn to the absence of an established basis for the reconstruction of past environments from insect death assemblages. A naïve reconstruction based on a modern death assemblage from a drain in the City of York is tested and shown to be inadequate in a number of respects. Certain of the habitats indicated by the fauna are absent from the surroundings, while the importance of some others is exaggerated. It is concluded that these errors were brought about by the active or passive long distance transport of insects.     

Insect and clitellate annelid traces in mesofossil assemblages from the Cretaceous of Australasia

Mesofossil assemblages from several Cretaceous and Cenozoic units across Australia and New Zealand provide new evidence of insect and annelid behaviour. The earliest scale insects (Diaspididae, Coccoidea) from Australasia are described and represented by three scale morphotypes. The mesofossil assemblages also reveal clitellate annelid cocoon morphotypes, three morphotypes of arthropod coprolites and several insect piercement structures on gymnosperm leaf or stem fragments, possibly related to feeding or more likely oviposition. This research offers a new avenue for detecting cryptic terrestrial invertebrate groups and their interactions, particularly with plants, in the fossil record. The fossils demonstrate that insect/invertebrate activity can be preserved and identified in mesofossil suites, that such traces and exoskeleton fragments are relatively common in acid-extracted mesofossil suites, and that recognizable categories occur on multiple landmasses and at various ages.     

The visibility of past trees and woodland: testing the value of insect remains

The reconstruction of woodland history is important in relation to archaeological, ecological, biogeographical and evolutionary problems, and insect remains are a significant source of relevant information. Fully natural interglacial and Holocene ‘waterlogged’ deposits assumed to have formed in woodland generally contain abundant macrofossils of both plants and insects indicative of trees. In contrast, British archaeological deposits rich in macrofossil remains of trees often lack, or contain very few, tree-associated insects. To cast light on this contradiction, assemblages of insect (Coleoptera and Hemiptera) remains from a range of modern deposits with various spatial relationships to woodland and trees have been analysed. The proportions of tree-associated insects varied greatly. There was a general trend from higher values in woodland and near to isolated trees of species supporting a rich insect fauna, to low or zero values where there were no trees. However, low values sometimes occurred in woods or near trees, so that rarity of tree-associated insects in archaeological deposits does not always carry the implication of a treeless environment. Further investigation is suggested, with emphasis on the importance of identifying isolated trees, scrub and hedges as a resource for humans and wildlife in the past.     

Isotopes as palaeoeconomic indicators: new applications in archaeoentomology

Previous studies have demonstrated the potential of archaeologically-recovered insect remains to stand as evidence of past human activities such as trade and industry. The present investigation reports the recovery of stable-isotopes from modern and Neolithic insect fauna and explores their utility in addressing issues of palaeoeconomic significance. While in its infancy, the application of stable-isotope methods towards archaeological insects may provide insight into exchange patterns and land-usage in the past. In the present analysis, stable-isotopes from modern and archaeological insects showed potential as a means of assessing local versus non-local signatures. Further study is recommended.     

A multivariate analysis of pottery from southwestern Cyprus using neutron activation analysis data

Neutron activation analysis is employed to geochemically characterize 129 pottery sherds collected from 38 sites during an archaeological survey of the Paphos District of southwestern Cyprus. Cluster analysis identifies three significant groupings using 26 trace elements and rare earth elements. These compositional groupings appear to reflect differences in the geographical locations of the sherd collection sites and particularly the influence of the surficial geology. In particular, the rocks of the ultramafic Troodos Ophiolite Zone, the Mamonia Complex, and various formations of the Circum Troodos Sedimentary Sequence appear to impart a detectable imprint on the sherds of the study area. Discriminant analysis identifies Ce, Th, Cr, Na, Fe, Hf, Ba and Co as the most significant chemical discriminators between the three compositional groupings and derives two discriminant functions that between them account for 100% of the variance between the groups. Discriminant analysis also confirms that 82% of the sherds classified by cluster analysis are correctly classified. Based on probability measures, the large majority of the sherds (98%) are considered to be local in origin.     

Insect invaders,seasonality and transhumant pastoralism in the Icelandic shieling economy

Abstract

The seasonal movement of people and animals to summer farms, or shielings in outfield pastures was a key element of Iceland's farming practice for over a millennium. At these sites, cattle and sheep husbandry, dairying and the harvesting of outfield resources took place. Despite their central role in the Icelandic economy, evidence for shielings in the landscape is ambiguous and the identification of a site as a shieling, as opposed to a farm, has relied upon written and place name evidence. The Norse colonists introduced a range of insects in their fodder, stored food, dunnage and ballast. Many of these are unable to live under natural conditions in Iceland and are dependent on people for survival. In 1991 Buckland and Sadler suggested that these species might be expected to be absent at shielings, as the sweepstake of their introduction and the seasonal hiatus in occupation would preclude their successful colonisation. This paper presents new evidence from a sub-fossil insect assemblage, which indicates that some of these insects are present at an Icelandic shieling. The implications of this for discerning the materials imported to shielings and the usefulness of Coleoptera for the identification of seasonality in the North Atlantic is discussed.     

Damage caused by insects during the mummification process: an experimental study

The process of mummification had been known since the Fourth Dynasty of the Old Kingdom (ca. 2600 bc) and continued to develop throughout ancient Egyptian history. Although mummification protected the body from decay, especially by microbes, some mummification techniques left the body susceptible to insect attack. Certain types of insects have been detected in the mummies. In scholarly publications, most authors have dealt with microorganisms, while few have concerned themselves in depth with the effect of insects on the mummies. This study aims to discuss the significance of insects and the changes they affected to the mummies during embalming. To achieve this goal, experiments were carried out replicating various mummification techniques using albino Wistar rats. Analysis and investigative techniques used included visual observation, Fourier transform infrared spectroscopy, investigation of the surface morphology by a scanning electron microscope, and color change by a spectrophotometer. The following insects could be identified as being present during the second and third processes of mummification: Dermestes maculatus, Necrobia rufipes, Saprinus gilvicornis, Chrysomya albiceps, Wohlfahrtia magnifica, and Attagenus fasciatus. In addition, the majority of our findings confirmed that the degradation by insects increased with the second and third methods of mummification. Finally, the experimental study conducted using the mummification techniques of the New Kingdom (ca. 1570–1070 bc) indicated that they were more resistant to insect attack than the other types.     

Insect Assemblages and Local Environment in the Mediaeval Town of Uppsala, Sweden

Insect remains from a mediaeval settlement in the town of Uppsala, S. Sweden, were analysed. Eighty-one insect taxa were identified from samples dating from the 12th to the 15th century. The insect assemblages are totally dominated by beetles. Only a few remains of butterflies, true flies and a bumble bee were found. The insects imply that the settlement was situated in open landscape. The settlement most likely consisted mainly of farm buildings throughout the studied period. Crops such as wheat, barley and cabbage were probably cultivated, particularly during the early settlement phases. Later, at the beginning of the 15th century, stock rearing seems to have dominated. The results suggest that the climate, during Mediaeval time in southern Sweden, was similar to the present or characterized by slightly higher summer temperatures. A number of currently very rare species were also recorded.     

Palaeoecology and palaeoenvironment of the Jurassic Talbragar Fossil Fish Bed,Gulgong, New South Wales,Australia

BEATTIE, R.G. & AVERY, S., December 2012. Palaeoecology and palaeoenvironment of the Jurassic Talbragar Fossil Fish Bed, Gulgong, New South Wales, Australia. Alcheringa 36, 451–465. ISSN 0311-5518.

The Talbragar Fossil Fish Bed has produced a significant number and variety of insect fossils, including the orders Hemiptera, Coleoptera, Orthoptera, Plecoptera, Odonata, Neuroptera, Diptera, Hymenoptera and Mecoptera. Hemiptera are the most common insects. Coleoptera are both common and diverse. Both orders include terrestrial and aquatic species. The other insect orders represented are less common. Many of the insect fossils discovered were previously unknown in the Australian region during the Jurassic, and the new records extend their palaeogeographic range into southeastern Gondwana. A recent collection of insects, fish, coprolites, gastropods, bivalves, bark, leaves, fruiting bodies, burrows and other ichnofossils supports a model of a shallow-water-lake palaeoenvironment at the northern end of the deposit, grading to a shoreline palaeoenvironment towards the southern end of the deposit. The existence of a productive lake supporting a large population of fish and a diverse, aquatic/shoreline and terrestrial woodland fringe palaeoecosystem, dominated by insects and woody plants, is demonstrated. There is no evidence of tetrapods. The Talbragar Fossil Fish Bed has similarities to Type A and B Mesozoic lake deposits reported from Transbaikalia. Culicimorph pupae are numerous and may have filled a significant niche near the base of the aquatic palaeoecosystem. Other immature aquatic insects are rare. The generally articulated nature of the fossil insects is interpreted to have resulted from a sudden but low-energy burial event.     

Dietary ecology of extant guanaco (Lama guanicoe) from Southern Patagonia: seasonal leaf browsing and its archaeological implications

The analysis of dietary traits of ungulates through tooth microwear and mesowear has been applied to archaeological sites to investigate seasonal changes in settlements by hunter–gatherers. In this paper we propose to test the hypothesis that tooth microwear (combined to mesowear) is able to indicate seasonality in the diet of extant ungulates in arid habitats (semi-deserts or steppe). The material analyzed comes from six faunal monospecific assemblages of guanaco (Lama guanicoe) resulting from a mass mortality event in winter 2000 near the Cardiel Lake in Southern Patagonia (Province of Santa Cruz, Argentina). Mesowear results indicate that the guanacos from the Cardiel Lake area are mixed feeders, and thus, have a diet that shifts seasonally. Moreover, microwear analysis supports the hypothesis that tooth microwear is able to indicate seasonality in the diet of extant guanaco in arid habitats. The pattern is clear for the winter sample and needs to be confirmed for a summer sample. Consequently, tooth microwear is proposed as a new potential proxy for detecting seasonal occupation in archaeological sites in Patagonia and other arid environments.     

To see or to be seen… is that the question? An evaluation of palaeolithic sites’ visual presence and their role in social organization

Visibility is one of the most common features considered when analyzing site location; visual control of a territory is usually considered to be a desirable characteristic for a prehistoric settlement. However, the role that visibility could have played in the perception of sites, as a significant element of palaeolithic landscapes, has been rarely evaluated. Sites can act not only as settlements but as places for population aggregations, social interactions and symbolic activities; in this context, the relevance presence of a site within the landscape could have been an influencing factor in site location preferences. This paper focuses on the visual presence of a set of sites from Late palaeolithic Cantabria (Northern Iberian Peninsula), and evaluates how the way they were perceived could have been related to their role in the social organization of foraging communities.     

Exaptation and synanthropic insects: A diachronic interplay between biology and culture

Abstract

Since 1982, Stephen J. Gould and Elisabeth Vrba's concept of exaptation has been adopted, to varying degrees, by a number of biological and social science disciplines. However, it remains a missing term in the study of archaeologically recovered insect remains and entomologically related artefacts. Exaptation refers to the process by which a characteristic acquires a function for which it was not originally adapted. The application of Gould and Vrba's concept to the field of archaeoentomology could enhance our perception of past social, environmental, and climatic conditions as well as interactions. Herein, biological and technological examples of the enduring relationship between synanthropic insects and humans are explored using the concept of exaptation.     

The application of numerical techniques to insect assemblages from the site of Stóraborg,Iceland

The problems of interpretation of many overlapping species lists of fossil assemblages are discussed using the example of insect faunas from a late medieval farm site in southern Iceland. Numerical techniques were employed, principally those available within the CLUSTAN package, and it was concluded that a considerable refinement in interpretation was achieved. Such quantitative procedures are suitable for groups other than fossil insects but their employment cannot be regarded as a substitute for, rather than an adjunct to, a sound ecological approach.     

Recovery of DNA from archaeological insect remains: first results,problems and potential

We report the recovery of short fragments of PCR amplifiable ancient DNA from exoskeletal fragments of the grain weevil Sitophilus granarius (L.) extracted from Roman and medieval deposits in Northern England. If DNA preservation in archaeological insect remains is widespread then many applications in the spheres of evolutionary studies and archaeology can be conceived, some of which are outlined.     

Aurignacian ethno-linguistic geography of Europe revealed by personal ornaments

Our knowledge of the migration routes of the first anatomically modern populations colonising the European territory at the beginning of the Upper Palaeolithic, of their degree of biological, linguistic, and cultural diversity, and of the nature of their contacts with local Neanderthals, is still vague. Ethnographic studies indicate that of the different components of the material culture that survive in the archaeological record, personal ornaments are among those that best reflect the ethno-linguistic diversity of human groups. The ethnic dimension of beadwork is conveyed through the use of distinct bead types as well as by particular combinations and arrangements on the body of bead types shared with one or more neighbouring groups. One would expect these variants to leave detectable traces in the archaeological record. To explore the potential of this approach, we recorded the occurrence of 157 bead types at 98 European Aurignacian sites. Seriation, correspondence, and GIS analyses of this database identify a definite cline sweeping counter-clockwise from the Northern Plains to the Eastern Alps via Western and Southern Europe through fourteen geographically cohesive sets of sites. The sets most distant from each other include Aurignacian sites from the Rhône valley, Italy, Greece and Austria on the one hand, and sites from Northern Europe, on the other. These two macro-sets do not share any bead types. Both are characterised by particular bead types and share personal ornaments with the intermediate macro-set, composed of sites from Western France, Spain, and Southern France. We argue that this pattern, which is not explained by chronological differences between sites or by differences in raw material availability, reflects the ethnolinguistic diversity of the earliest Upper Palaeolithic populations of Europe.