A short note on starch and xylem of Colocasia esculenta (taro) in archaeological deposits from Pitcairn Island,southeast Polynesia

We present results of starch analysis of archaeological deposits from Pitcairn Island. High concentrations of starch grains preserved in cell membranes, and xylem tracheary elements, consistent with introduced Colocasia esculenta (taro) were found. Because of limited age control, we are uncertain if the microfossils are prehistoric. Problems associated with identifying taxa with small starch grains in extractions from weathered deposits are highlighted.     

Archaeobotanical evidence of prehistoric maize (Zea mays) consumption at the northern edge of the Great Plains

Analysis of starch granules, phytoliths, and plant macrofossils from archaeological features and carbonized food residue provides important new insight into the extent of prehistoric maize (Zea mays) consumption on the North American Great Plains. These data suggest that consumption of maize, and probably other cultigens, was widespread on the eastern Canadian Prairies between approximately AD 1000 and 1600. Domesticated plants may have been grown locally, acquired through trade, or transported into the region following dispersal of family groups from horticultural villages located elsewhere. However, the lack of strong artifactual evidence of gardening, and the small-scale nature of sites on the eastern Canadian Prairies indicate that local horticulture, if practiced, was non-intensive.     

Plant microfossil analysis reveals disturbed forest and a mixed-crop,dryland production system at Te Niu,Easter Island

Plant microfossil analysis was carried out on 12 soil samples from a variety of landscape features at three locations (100–250 m a.s.l.) along a 950 m transect at Te Niu, Easter Island. Pollen and phytolith assemblages were dominated by palms and ground ferns, and suggested disturbed forest. We identified pollen of bottle gourd (Lagenaria siceraria) and starch grains of the common yam (Dioscorea alata), sweet potato (Ipomoea batatas) and taro (Colocasia esculenta). The data suggest a mixed-crop, dryland production system at Te Niu dominated by yam and sweet potato, and supplemented by taro and bottle gourd. The data provide direct evidence of crop type and range, supporting the indirect evidence (topographic and landscape features, field and historical research, comparisons with elsewhere in the Pacific) that much of the rock covered landscape of Easter Island was used for intensive horticulture.     

Plant microfossils in prehistoric archaeological deposits from Yuku rock shelter,Western Highlands,Papua New Guinea

Plant microfossil analysis was carried out on anthropogenic deposits from Yuku rock shelter, possibly occupied from before 14,200 cal. BP to recent times, in the Papua New Guinea Highlands. Elaeocarpus pollen has extremely high percentages throughout. Starch residues of lesser yam (Dioscorea esculenta) and one or more, undifferentiated Dioscorea species were identified, suggesting processing of these taxa since first use of the site. The latter include Dioscorea alata, Dioscorea bulbifera, Dioscorea nummularia and Dioscorea pentaphylla. Pollen and phytoliths of banana (Musa) appear 5200 cal. BP. We cannot differentiate between naturally growing and cultivated yams and bananas. Putative truffle (hypogeous Ascomycotina) spores in the deposits suggest foraging of truffles.     

Boat-based foraging and discontinuous prehistoric red abalone exploitation along the California coast

Temporally and spatially discontinuous pulses of heavy prehistoric exploitation of red abalone (Haliotis rufescens) have been documented along the southern and central California coast. This article examines the very late (post-950 cal BP) appearance of numerous red abalone processing sites on the Monterey Peninsula in central California. We test three prominent explanations offered for the sudden onset of red abalone processing sites: trophic cascades resulting from human predation on sea otters, logistical foraging by inland residents, and changes in sea surface temperature. A trophic cascade appears to have occurred but does not fully explain the nature or timing of the phenomenon in the region.     

Red abalone collecting and marine water temperature during the Middle Holocene occupation of Santa Cruz Island,California

Oxygen isotope values derived from prehistoric mussel (Mytilus californianus) shell calcite are used to determine whether sea-surface temperatures in the vicinity of the Punta Arena site (CA-SCRI-109) on Santa Cruz Island, California, were cooler than present between 6300 and 5300 cal BP. This site and others in the western sector of the island dating to this period are distinctive because of the presence of large red abalone (Haliotis rufescens) shells, a species that historically has been subtidal around Santa Cruz and the other northern Channel Islands. Comparison of temperature values derived from archaeological shells with those from modern mussel shells, along with water temperatures derived from satellites, indicates that waters were cooler during the 6300 to 5300 cal BP period. These results are consistent with a previous study and support the interpretation that collection of red abalone during the 6300–5300 cal BP period was partly the result of cooler water temperatures that made this large mollusk more available to foragers in the intertidal or shallow subtidal zone.     

A short note on banana (Musa) phytoliths in Lapita,immediately post-Lapita and modern period archaeological deposits from Vanuatu

We present results of phytolith analysis of deposits from three prehistoric settlement sites in Vanuatu: the small islands of Vao and Uripiv off the north east coast of the main island of Malekula and at Mafilau on the west coast of Epi Island. Samples from Vao Island coarsely encompass the period from Lapita times (c. 3000 BP) to the present. The data from this site suggest deforestation by people, with a decrease in palms and increase in grasses. The data from the three sites show evidence of an introduced cultigen during different prehistoric periods. Banana (Musa) phytoliths were found in one recent Vao layer (500 BP-present) and in Mafilau layers, the latter representing the immediate post-Lapita period (c. 2800–2500 BP). This phytolith type was also found at Uripiv in Lapita layers (c. 3000–2700 BP). Banana phytoliths are diagnostic so their presence in archaeological deposits in Remote Oceania, outside their natural distribution, provides secure evidence of cultivation. This demonstrates the potential of this line of evidence for identifying both the cultigens transported by early colonists and subsequent plant introductions. The data provide the first direct evidence of banana cultivation at a Lapita site in Remote Oceania and support the contention that horticulture was an integral part of the Lapita Cultural Complex.     

Starch grain characterization of Prosopis chilensis (Mol.) Stuntz and P. flexuosa DC,and the analysis of their archaeological remains in Andean South America

The fruit (pods) of Prosopis (Fabaceae) are frequently recovered from pre-Hispanic Argentinian archaeological sites, suggesting that this genus was of importance in ancient economies in this region. Yet it is only recently that archaeobotanists have begun to carry out systematic research into this genus. Therefore many questions remain to be addressed concerning the food value of Prosopis fruit, and its potential contribution. This paper examines starch from the pods of two species, Prosopis flexuosa and Prosopis chilensis, for the purposes of describing and classifying their morphological features and biometrical parameters. Pods of both species were gathered from two extremes (northern and southern) of the Hualfín Valley, Catamarca, Argentina. Starch abundance is estimated and compared with that of Zea mays, an economically important plant with high concentrations of starch. This paper reveals that Prosopis pods contain very low amounts of starch compared to starch-rich edible species. Despite this, the recovery of Prosopis starch grains can be successfully used to assess archaeological tool uses. Prosopis starch grains were found to be highly variable in shape. Grain size ranged between 10 and 20 μm. Granule irregularity and the high birefringence under polarized light are two of the most important diagnostic features. No significant statistical differences were found in the structure (morphology, size and hilum) of the starch of the same species from both localities. Finally, the implications for the role of this plant in past societies are discussed.     

Starch grains analysis of stone knives from Changning site,Qinghai Province,Northwest China

Plant residues recovered from prehistoric stone artifacts can be used to help explain tool function and plant use. At the Changning site in Qinghai Province, Northwest China, dating from 4000 yr BP, we examined starch granules extracted from three slate stone knives. A total of 153 starch grains were retrieved from three stone knives, from which we identified starches from legumes, the Triticeae tribe, foxtail millet (Setaria italica), broomcorn millet (Panicum miliaceum), roots and tubers. These results indicate that the stone knives may have been used for a variety of activities that included reaping grasses and food processing. The species of starch grains retrieved from the study sample reveal that diverse crops were cultivated at the Changning site 4000 years ago.     

Irrigated taro (Colocasia esculenta) farming in North Kohala,Hawai‘i: sedimentology and soil nutrient analyses

How did traditional farming transform the natural environment in the Hawaiian Islands? This question is one that has largely been addressed for rainfed farming of crops like sweet potato (Ipomoea batatas), but evidence is lacking for irrigated farming of the critical staple crop taro (Colocasia esculenta). We describe the results of soil nutrient and sedimentological analyses of deeply-stratified pondfield deposits representing a 600-year-long record of irrigated taro farming in the North Kohala District, Hawai‘i Island. Soil is categorized by particle size to determine modes of transport and deposition, and concurrent soil nutrient analyses were conducted to infer shifts in the source of sediments and changes associated with taro harvesting. The advent of farming is clearly detectable in sedimentology, the presence of charcoal found within sediments, and soil chemistry. However, diminished nutrient concentrations can be attributed largely to deposition of a mixture of upstream sediments. Overall, there is no clear evidence for nutrient draw-down by taro harvesting, but we cannot yet rule it out as a factor. This study demonstrates the inherent difficulty of correlating changes in soil nutrients evident in irrigated pondfields with the long term history of soil nutrient cycling.     

Sweet potato (Ipomoea batatas) and banana (Musa sp.) microfossils in deposits from the Kona Field System,Island of Hawaii

Analysis of plant microfossils (pollen, phytoliths, starch grains and xylem cells) was carried out on 12 soil samples from a variety of pre-contact archaeological features in South Kona within a portion of what has been termed the Kona Field System, on the Island of Hawaii. The oldest radiocarbon ages of the sampled deposits are 1300–1625 AD and 1310–1470 AD. The pollen and phytolith evidence suggests a change from more to less trees and shrubs in the area as a result of human activity. We found phytoliths of banana (Musa sp.) leaves in most and starch grains and xylem cells of tuberous roots of c.f. sweet potato (Ipomoea batatas) in all of the samples, suggesting that the sampled features were associated with these crops and that both crops were cultivated intensively within the study area. Higher concentrations (volumetric) of starch and xylem in samples from older deposits suggest that cultivation was more intensive then. The apparent absence of starch and xylem remains of other tuberous crops archaeologically identified elsewhere in Polynesia suggests that tuberous cropping within the study area was mono-specific.     

Prehistoric behavioral depression of cormorant (Phalacrocorax spp.) on the northern California coast

Resource depression is often assumed to result from the over-exploitation of a prey species leading to declines in its population, and therefore availability, to human hunters. However, two other forms of resource depression are noted in the ecological literature: behavioral and micro-habitat depression. Data presented in this paper demonstrate resource depression of cormorants (Phalacrocorax spp.), a coastally breeding and semi-pelagic bird genus, at a coastal site in northern California. Citing modern cormorant behavior and a lack of evidence for intensive rookery exploitation, it is argued that the observed resource depression resulted from changes in cormorant behavior that decreased encounter rates for human hunters. Such behavioral depression is cited as a buffer to the extirpation of prey in mainland settings.     

Disentangling long bones of foxes (Vulpes vulpes and Alopex lagopus) from arctic archaeological sites

To discern the presence of two anatomically close species within an archaeological assemblage is always a problem for the analyst. A particularly interesting case is the distinction between the red fox (Vulpes vulpes) and the arctic fox (Alopex lagopus), which is often determined using non-rigorous criteria and producing results that must be regarded with caution. Classical methods for separating these two species and determining sexual dimorphism, and eventually the sex ratio, such as analyses using uni- and bivariate plots, are based on arbitrarily fixed limits between sexes. In this text, a more robust statistical method termed mixture analysis (MA) is used to determine the species of limb bones from foxes. First, the MA is applied to a sample of each species using metric data taken from a collection of present-day fox skeletons. Afterwards, the MA is applied to archaeological samples dated to the Dorset period and retrieved from the Tayara site (near Salluit, South Hudson Strait, Nunavik). The results clearly demonstrate that the greatest length (GL) of long bones, especially the humerus and tibia, is the best measurement for distinguishing the species, followed by the distal breadth and the proximal breadth; and that GL is better suited for sexing foxes. The results of MA applied to the Tayara site collection lead to the conclusion that the red fox is present along with the arctic fox among the assemblage, and that there is no fox body size change during the last two millennia in the Eastern Arctic.     

Morphometric and taphonomic analysis of granular ark (Anadara granosa) dominated shell deposits of Blue Mud Bay,northern Australia

Metrical analyses of complete mollusc shells have commonly been used to examine aspects of prehistoric mollusc demography, and by extension to investigate potential human exploitation of a variety of species. Recently it has been highlighted that several taphonomic processes, such as fragmentation, have the potential to introduce size bias into these archaeological samples, particularly towards smaller size classes, and thus the potential to significantly skew results of these metrical analyses. This is investigated here by deriving morphometric equations for the sand/mudflat bivalve Anadara granosa, a species prevalent in archaeological sites across northern Australia. A linear regression model based on a series of valve measurements, resulting in high coefficients of determination, demonstrates strong predictive power for size reconstruction. The effects of fragmentation on assemblages from two case study sites are then examined as a means of reviewing issues of differential size preservation, variation in mean valve size, and evaluating the strength of previous interpretations of intensive human exploitation in Blue Mud Bay. Based on these analyses it appears that A. granosa mean size is not significantly affected by fragmentation as other species described in the literature, and previous interpretations retain their validity.     

Aging mountain gazelle (Gazella gazella): refining methods of tooth eruption and wear and bone fusion

This paper refines current methods for aging mountain gazelle (Gazella gazella), one of the most common species hunted by prehistoric humans in the southern Levant. Most importantly, we present a new tooth wear scheme for aging the lower deciduous fourth premolar (dP₄), the lower permanent molars (M₁, M₂, M₃) and the lower fourth premolar (P₄) of the gazelle. Broader wear stages for complete mandibles and individual teeth and bone fusion stages are also provided. Finally, a comparison of the tooth wear and bone fusion data indicates that the two methods can be linked at the important transition when a gazelle reaches adulthood (ca. 18 months of age). The completion of the fusion of the gazelle skeleton corresponds directly to the replacement of the dP₄ by the P₄ and the beginning of wear of the M₃. The coincidence of these aging indicators allows for direct comparison of gazelle bone fusion and tooth wear data.     

Starch analysis reveals prehistoric plant translocations and shell tool use,Marquesas Islands,Polynesia

Starch analysis is proving particularly useful in tropical regions like the central Pacific where crop inventories are often dominated by starchy fruits and tubers and recovery of macrobotanical remains is rare. Analysis of 23 shell tools from the Marquesas Islands provides direct evidence for translocation of five traditional crop plants. Four taxa derive from the western Pacific, including Artocarpus altilis (breadfruit), Piper methysticum (kava), Colocasia esculenta (taro), and one or more species of Dioscorea (yam). The fifth taxon, the South American Ipomoea batatas (sweet potato), dates from the 14th century AD onward and constitutes the earliest record of this cultigen in the archipelago. The occurrence of sweet potato starch suggests that this crop plant may have been more important than usually is assumed, while the limited recovery of breadfruit starch, the main Marquesan food plant at western contact, requires further investigation. The starch residues also inform on tool use, demonstrating that shell tools ethnographically associated with specific crops (e.g., “breadfruit peelers”) had more generalised functions. This study is a further demonstration of the potential of starch analysis to provide important information on the history of crop introductions, on-site activities, and artefact use.     

Focused,intense and long-term: evidence for granular ark (Anadara granosa) exploitation from late Holocene shell mounds of Blue Mud Bay,northern Australia

Explanatory frameworks relating to the appearance of shell mounds and the exploitation of molluscs (particularly the sand/mudflat bivalve Anadara granosa) during the late Holocene have tended to emphasise stasis and continuity. Very few analyses have adequately investigated the intensity of human predation during the mound period and the potential effects on the particular prey species. To this end, the biological and ecological characteristics of A. granosa, the dominant molluscan species for much of the known period of occupation in the region, are considered in detail, in combination with assessing the potential for human impact through predation via the measurement of 9106 A. granosa valves from three shell mounds. In explaining long-term economic change in this region of northern Australia, the focus has been placed on the analysis of relative changes and trends through time in prehistoric resource exploitation.     

Introduced taro (Colocasia esculenta) and yams (Dioscorea spp.) in Podtanean (2700–1800 years BP) deposits from Mé Auré Cave (WMD007), Moindou,New Caledonia

Plant microfossil analysis was carried out on deposits bearing Podtanean (2700–1800 years BP) ceramics from Mé Auré Cave, Moindou, New Caledonia. Starch residues of introduced Colocasia esculenta, Dioscorea esculenta and one or more, undifferentiated Dioscorea species were identified. The latter include Dioscorea alata, D. bulbifera, D. nummularia and D. pentaphylla. The data provide the first direct evidence of prehistoric horticulture in New Caledonia.     

A deciduous and permanent dental wear stage system for assessing the age of Trachypithecus sp. specimens (Colobinae, Primates)

Although Colobine monkeys are one of the most common prey hunted by Southeast Asian prehistoric humans, no data concerning tooth eruption and wear are available for these species. Dental wear eruption and attrition are used for attributing ages to individual fossils in order to construct mortality curves, and are also useful in reconstructing fossil life histories. Such wear sequences partly exist for Cercopithecines. Although Cercopithecines and Colobines are both bilophodont, they present significant differences in terms of diet, detailed tooth morphologies and occlusion. This paper aims to formulate a guide for the dental eruption and attrition of extant Trachypithecus specimens, the most folivorous of all the Colobines. We also propose to calibrate the eruption and wear stages that we define here with absolute ages available in the literature.