Modelling the effect of Mesolithic populations on the slowdown of the Neolithic transition

The expansion of the Neolithic transition in Europe took place gradually from the Near East across the whole continent. At Northern Europe, observations show a slowdown in the speed of the Neolithic front in comparison to other regions of the continent. It has been suggested that the presence of high population densities of hunter-gatherers at the North could have been the main cause for this slowdown. This proposal has recently been described by a mathematical model that takes into account: (i) the resistance opposed by the Mesolithic populations to the advance of Neolithic populations in their territory, and (ii) a limitation on the population growth dynamics due to the competition for space and resources. But these two effects are not equally responsible for the slowdown of the spread. Indeed, here we show that the limitation on the population growth dynamics seems to have been the main cause of the delay of the expansion of farming in Northern Europe.     

A Variation on a Basic Theme: The Transition to Farming in Southern Central Europe

This paper attempts to summarize the past years of research on the Mesolithic–Neolithic transition in Central Europe and to review recent discussions about the origin and spread of the Early Neolithic. Particular emphasis is given to the debate about migration or diffusion. A combined migrationist/diffusionist model is presented, arguing for an emergence of a farming economy among hunter-gatherer populations in Transdanubia and the subsequent spread of this economy through migration. The new settlers interacted with local Mesolithic groups and adopted and incorporated local material culture and sometimes even aspects of local Mesolithic economy, a process which continued throughout the Early Neolithic. With time, population increase, subsequent competition for resources, and climatic instability led to a destabilization of traditional Early Neolithic society and finally to the outbreak of severe intercommunity violence. The only escape from mutual extinction was a rearrangement of subsistence and social and political structures, possibly with contributions from surviving Terminal Mesolithic groups.     

Prehistoric population history: from the Late Glacial to the Late Neolithic in Central and Northern Europe

Summed probability distributions of radiocarbon dates are used to make inferences about the history of population fluctuations from the Mesolithic to the late Neolithic for three countries in central and northern Europe: Germany, Poland and Denmark. Two different methods of summing the dates produce very similar overall patterns. The validity of the aggregate patterns is supported by a number of regional studies based on other lines of evidence. The dramatic rise in population associated with the arrival of farming in these areas that is visible in the date distributions is not surprising. Much more unexpected are the fluctuations during the course of the Neolithic, and especially the indications of a drop in population at the end of the LBK early Neolithic that lasted for nearly a millennium. Possible reasons for the pattern are discussed.     

Neolithic economy in central Europe

This article presents a survey of Neolithic economy, land use, trade, natural environment, and usage of plant and animal resources in central Europe, 5415–2240 B.C. (4500–1800 bc). Early, Middle A and B and Late Neolithic materials are summarized and compared. The earliest farmers expanded from southern Hungary and adjacent areas into central Europe over a relatively short time period, 100–200 years. They occupied areas only with good soils; thus probably hunters and gatherers continued to exist in many regions of central Europe. There is an increase in population and more upland areas are exploited for farming during the Middle Neolithic A and B periods. By the Middle Neolithic B period, low-level hierarchical or ranked societies appear in some regions of central Europe. The Late Neolithic may represent a modification of the mixed farming strategy observed during the earlier periods. Perhaps the herding of domestic animals became more important.     

Spatial dimensions increase the effect of cultural drift

The transition from hunter-gathering to agriculture (Neolithic) spread gradually across Europe from the Southeast. A reduction in cultural diversity of crop farming practices has been previously observed by comparing pre-LBK Neolithic sites in Greece and the Balkans (dated about 8500 yr BP) to LBK Neolithic sites in Central Europe (dated about 7000 yr BP). The decrease in crop diversity is statistically significant even when considering only the species less likely to have been subject to smaller productivity due to climatic factors (reductions in growing season, temperature, daylight, etc.). This reduction in cultural diversity has not been explained previously. In this paper we show that spatial drift, which occurred on the front of the advancing wave of pioneer settlements, can explain the observed loss of diversity during the LBK range expansion. Our results suggest that spatial dimensions can have a relevant effect also in other case studies in which cultural drift is important.     

Ecological constraints on the first prehistoric farmers in Europe

The Neolithic Revolution, which witnessed the transformation of hunter–gatherer groups into farming communities, is traditionally viewed as the event that allowed human groups to create systems of production that, in the long run, led to present-day societies. Despite the large corpus of research focused on the mechanisms and outcomes of the Neolithic transition, relatively little effort has been devoted to evaluating whether particular production-oriented adaptations could be integrated into a broad range of ecological conditions, and if specific cultural traditions differed ecologically. In order to investigate whether the differences between the adaptations and geographic distributions of three major Early Neolithic archaeological cultures are related to the exploitation of different suites of environmental conditions, we apply genetic algorithm and maximum entropy ecological niche modeling techniques to reconstruct and compare the ecological niches within which three principal Neolithic cultures (Impressed Ware, Cardial Ware, and Linearbandkeramik) spread across Europe between ca. 8000 and 7000 cal yr BP. Results show that these cultures occupied mutually exclusive suites of environmental conditions and, thus, were adapted to distinct and essentially non-overlapping ecological niches. We argue that the historical processes behind the Neolithization of Europe were influenced by environmental factors predisposing occupation of regions most suited to specific cultural adaptations.     

Ancient DNA insights from the Middle Neolithic in Germany

Genetic studies of Neolithic groups in central Europe have provided insights into the demographic processes that have occurred during the initial transition to agriculture as well as in later Neolithic contexts. While distinct genetic patterns between indigenous hunter-gatherers and Neolithic farmers in Europe have been observed, it is still under discussion how the genetic diversity changed during the 5,000-year span of the Neolithic period. In order to investigate genetic patterns after the earliest farming communities, we carried out an ancient mitochondrial DNA (mtDNA) analysis of 34 individuals from Wittmar, Germany representing three different Neolithic farming groups (ca. 5,200–4,300 cal bc) including Rössen societies. Ancient DNA analysis was successful for six individuals associated with the Middle Neolithic Rössen and observed haplotypes were assigned to mtDNA haplogroups H5, HV0, U5, and K. Our results offer perspectives on the genetic composition of individuals associated with the Rössen culture at Wittmar and permit insights into genetic landscapes in central Europe at a time when regional groups first emerged during the Middle Neolithic.     

A Diachronic Study of Networks of Ceramic Assemblage Similarity in Neolithic Western Anatolia,the Aegean and the Balkans (c.6600–5500 bc)

This study compares the results of Jaccard and Kulczynski‐2 similarity measures on a sample of ceramic assemblages to reveal spatio‐temporal patterns in the relationship between Neolithic sites in western Anatolia and south‐eastern Europe (c.6600–5500 bc ). The results show that the relationship between spatial distance and ceramic assemblage similarity has increased through time, which supports previous interpretations of leapfrog migrations and subsequent regionalization in ceramic assemblages during the Neolithic. A diachronic network analysis demonstrates the continuation of Aegean networking after the spread of farming.     

Mobility and Social Change: Understanding the European Neolithic Period after the Archaeogenetic Revolution

This paper discusses and synthesizes the consequences of the archaeogenetic revolution to our understanding of mobility and social change during the Neolithic period in Europe (6500–2000 BC). In spite of major obstacles to a productive integration of archaeological and anthropological knowledge with ancient DNA data, larger changes in the European gene pool are detected and taken as indications for large-scale migrations during two major periods: the Early Neolithic expansion into Europe (6500–4000 BC) and the third millennium BC “steppe migration.” Rather than massive migration events, I argue that both major genetic turnovers are better understood in terms of small-scale mobility and human movement in systems of population circulation, social fission and fusion of communities, and translocal interaction, which together add up to a large-scale signal. At the same time, I argue that both upticks in mobility are initiated by the two most consequential social transformations that took place in Eurasia, namely the emergence of farming, animal husbandry, and sedentary village life during the Neolithic revolution and the emergence of systems of centralized political organization during the process of urbanization and early state formation in southwest Asia.

     

Founder effect, drift, and adaptive change in domestic crop use in early Neolithic Europe

We document and quantify a significant reduction in crop diversity in the early central European Neolithic using a large multi-site database of archaeobotantical remains we compiled from published Neolithic sites across southwest Asia and Europe. Two hypotheses are proposed to account for the observed changes: one which claims that the different environmental conditions of central Europe selected for a different set of crop choices and strategies than in use in southeast and Mediterranean Europe; and a null hypothesis that explains the change as a drift process associated with a small founding population that subsequently undergoes rapid expansion. Through an agent-based simulation model, we test the null hypothesis and demonstrate that the drop in diversity exceeds that predicted by a drift process. We conclude by re-evaluating the possible adaptive changes underlying crop use in early Neolithic Europe.     

Detection of diffusion and contact zones of early farming in Europe from the space-time distribution of 14C dates

The spread of early farming in Europe is revisited using a sample of 3072 audited 14C calBC dates from 940 georeferenced early Neolithic sites. The surface expansion of early Neolithic has been reconstituted using the kriging technique of spatial interpolation. Centres of renewed expansion, of contact zones, and the main routes of expansion have been highlighted by means of a vector map, representing the gradient. The expansion of the agricultural system on the map, was not uniform and regular across Europe as a whole, but proceeded in leaps. With the scale of detection of the 500-year isochrones, several leaps are identifiable: at 8000 calBC crossing the Taurus barrier, 6700–6100 calBC crossing the southern Adriatic barrier, 6100–5600 calBC crossing the Central European agro-ecological barrier and 5000–4000 calBC expanding on the other, marginal zones. Using a vector map, 10 points of renewed expansion and nine contact zones, were detected. The whole does not correspond to a process of homogeneous diffusion, approximately steady, but a process marked by phases of geographical expansion and stasis.     

A simulation of the Neolithic transition in Western Eurasia

Farming and herding were introduced to Europe from the Near East and Anatolia; there are, however, considerable arguments about the mechanisms of this transition. Were it the people who moved and either outplaced or admixed with the indigenous hunter-gatherer groups? Or was it material and information that moved—the Neolithic Package—consisting of domesticated plants and animals and the knowledge of their use? The latter process is commonly referred to as cultural diffusion and the former as demic diffusion. Despite continuous and partly combined efforts by archaeologists, anthropologists, linguists, palaeontologists and geneticists, a final resolution of the debate has not yet been reached. In the present contribution we interpret results from the Global Land Use and technological Evolution Simulator (GLUES). This mathematical model simulates regional sociocultural development embedded in the geoenvironmental context during the Holocene. We demonstrate that the model is able to realistically hindcast the expansion speed and the inhomogeneous space-time evolution of the transition to agropastoralism in western Eurasia. In contrast to models that do not resolve endogenous sociocultural dynamics, our model describes and explains how and why the Neolithic advanced in stages. We uncouple the mechanisms of migration and information exchange and also of migration and the spread of agropastoralism. We find that (1) an indigenous form of agropastoralism could well have arisen in certain Mediterranean landscapes but not in northern and central Europe, where it depended on imported technology and material; (2) both demic diffusion by migration and cultural diffusion by trade may explain the western European transition equally well; (3) migrating farmers apparently contribute less than local adopters to the establishment of agropastoralism. Our study thus underlines the importance of adoption of introduced technologies and economies by resident foragers.     

The Origins of the Neolithic Along the Atlantic Coast of Continental Europe: A Survey

The main aim of this work is to compare the processes of transition to the Neolithic along the Atlantic coasts of continental Europe. Archaeological data on the late Mesolithic and the early Neolithic in the best known regions (central and southern Portugal, Cantabrian Spain, Atlantic France, the shores of the North Sea, and southern Scandinavia) are discussed. The transition to the Neolithic in Atlantic Europe can be viewed as a relatively late phenomenon, with several interesting particularities. Among those, we point out the fundamentally indigenous character of the processes; the existence of a long availability phase, in which hunter-gatherer groups maintained contact with neighboring agriculturalists and probably were familiar with farming and animal husbandry without applying them in a systematic way; and the later development of megalithic monumental funerary architecture. Finally, the main hypotheses so far proposed to explain the change are contrasted with the available evidence: those that argue that the change derives from economic disequilibrium, and those that opt for the development of social inequality as the fundamental cause.     

The chronology of Neolithic dispersal in Central and Eastern Europe

We analyze statistically representative samples of radiocarbon dates from key Early Neolithic sites in Central Europe belonging to the Linear Pottery Ceramic Culture (LBK), and of pottery-bearing cultures on East European Plain (Yelshanian, Rakushechnyi Yar, Buh-Dniestrian, Serteya and boreal East European Plain). The dates from the LBK sites form a statistically homogeneous set with the probability distribution similar to a single-date Gaussian curve. This implies that the duration of the spread of the LBK is shorter than the available temporal resolution of the radiocarbon dating; therefore, the rate of spread must be larger than 4 km/yr, in agreement with earlier estimates. The East European sites exhibit a broad probability distribution of dates. We identify in these data a spatio-temporal sequence from south-east to north-west, which implies the rate of spread of the initial pottery-making of the order of 1.6 km/yr, comparable to the average rate of spread of the Neolithic in Western and Central Europe. We argue that this spatio-temporal sequence is consistent with an idea that the tradition of the initial pottery-making on East European Plain developed under an early impulse from the Eastern Steppe.     

The spread of Neolithic plant economies from the Near East to northwest Europe: a phylogenetic analysis

Phylogenetic techniques are used to analyse the spread of Neolithic plant economies from the Near East to northwest Europe as a branching process from a founding ancestor. The analyses are based on a database of c. 7500 records of plant taxa from 250 sites dated to the early Neolithic of the region in which they occur, aggregated into a number of regional groups. The analysis demonstrates that a phylogenetic signal exists in the data but it is complicated by the fact that in comparison with the changes that occurred when the crop agriculture complex expanded out of the Near East, once it arrived in Europe it underwent only limited further changes. On the basis of the analysis it has been possible to identify the species losses and gains that occurred as the complex of crops and associated weeds spread and to show the influence of geographical location and cultural affinity on the pattern of losses and gains. This has led to consideration of the processes producing that history, including some reasons why the dispersal process did not produce a perfect tree phylogeny, as well as to the identification of some specific anomalies, such as the unusual nature of the Bulgarian pattern, which raise further questions for the future.     

Regional Demographic Dynamics in the Neolithic Transition in Iberia: Results from Summed Calibrated Date Analysis

This paper assesses long-term trends in human activity across the Iberian Peninsula associated with the spread of agriculture. In order to improve our understanding of regional trajectories within the two-way and multiple-velocity model proposed by Isern et al. (Journal of Archaeological Method and Theory 21:447–460, 2014), we consider summed calibrated radiocarbon dates of a large audited dataset from across Iberia. To assess the role of different analytical units, two alternative peninsular divisions at different spatial resolutions were used: one, with three generalized study areas, and the second, with eight biogeographic regions. This comparison of spatio-temporal variability provides a more in-depth inter-regional reading of the Neolithic transition in Iberia than more generalized approaches. The southwest, Mediterranean coasts and interior portions of Iberia all showed significant demographic expansion with the onset of the Neolithic, as observed across Western Europe. By contrast, northern Iberia featured a close adherence to a null model of exponential growth and a longer continuation of hunter-gatherer strategies.     

Clearances and Clearings: Deforestation in Mesolithic/Neolithic Britain

Clearances, interpreted from pollen records during the Mesolithic and Neolithic of Europe, are generally ascribed to purposive deforestation which is compatible with the transition model, whereby early Neolithic economic strategies are a development of late Mesolithic intensification of wild plant food husbandry. This paper considers the role of natural processes in creating clearings and the role of inadvertent impact of human activity on forest processes, including woodland regeneration. The role of climate, wind-throw and lightning strikes in creating clearings and forest instability is emphasised and the evidence discussed from sites which may be interpreted as resulting from opportunistic human use of natural clearings. Unfortunately, regional pollen diagrams lack sufficient spatial resolution to detect the size of isolated clearings or establish the spatial variation in forest composition that was intimately related both to forest ecology and the effects of subtle human impacts. This may be the major reason for an apparent contradiction between pollen evidence of Neolithic impact and the archaeological record. Moreover, early Neolithic agricultural activity may have been concentrated in valley bottoms, which is undetectable in regional pollen diagrams. Alternative models need to be considered, which include culturally specific exploitation of the local environment, along with the inadvertent ecological repercussions of pre-agricultural and early-agricultural human activities in naturally dynamic woodlands.     

How was Bell Beaker economy related to Corded Ware and Early Bronze Age lifestyles? Archaeological,botanical and palynological evidence from the Hegau,Western Lake Constance region

Abstract

The Europe wide spread of what has been called the Bell Beaker phenomenon remains an enigma of European prehistory. While most of the recent research stresses the ideological aspects of using Bell Beaker material culture, here we take a regional and economical perspective. We look for the chronological relationships and the economic choices of the Bell Beaker phase and of its closest neighbours in time and space: the Late Neolithic Corded Ware and the Early Bronze Age. We focus on the regional archaeological settlement history and present the hitherto richest European Bell Beaker-associated collection of palaeobotanical macro-remains, together with our high-resolution palynological work on annually laminated lake sediments. These different lines of evidence are tied together by an absolute chronology derived from new radiocarbon accelerated mass spectrometry (AMS) dates (now more than 200) and from the dendrodates from the World Heritage wet preserved pile dwellings. We show the preceding Late Neolithic, the actual Bell Beaker, and the following Early Bronze age economies each relying on different agricultural strategies that focus on distinct parts of the landscape. There is no link obvious between Late Neolithic and Bell Beaker, but there is between Bell Beaker and Early Bronze Age. Related to different modes of production, differences in ideology become visible in food preferences as well as in other parts of the material culture. We conclude that the Bell Beaker economy represents a re-orientation of the mode of production focusing on single, rather small farmsteads which often do not leave a distinct signal in the archaeological record.     

The Italian Neolithic: A Synthesis of Research

This paper reviews the distinctive economic and social development of Neolithic Italy and its islands, from ca. 6000 B.C. until the emergence of the Copper Age ca. 3500 B.C. Through a synthesis of traditional interpretations and more recent discussions relating to early farming economies, social and technological developments, settlement, and landscape, the complex regional patterns are described. The development of archaeological studies in Italy, the biases in regional and chronological data collection, the regional patterns of cultures and landscapes, and the emergence of distinctive funerary, artistic and economic activities reveal a wealth of varied and intriguing archaeological information from a wide variety of sources that link parallel developments in the Mediterranean and Europe.     

THE FORMATION OF SOCIAL RANK IN THE EARLY NEOLITHIC OF NORTHERN EUROPE

Klavs Randsborg has made important contributions to the archaeology of the Bronze Age and later prehistoric periods, but in the 1970s he also touched upon issues such as the formation of rank in Neolithic societies. In his article ‘Social Dimensions of Early Neolithic Denmark’, he suggested that a hierarchical society arose at the transition from the Early to Middle Neolithic, c. 3300–3200 BC. Since then, excavations and research have resulted in numerous publications about the Neolithic, but only rarely have these examined social development. In this article, the authors continue the debate, sharing the same starting point as Klavs Randsborg, but here approaching the question of emerging social inequality on the background of recent research into the early agricultural societies in Northern Europe, seen in a broader European context. The primary focus is upon burial monuments as manifestations of status and power, and parallels are drawn with similar construction activities amongst present‐day farming communities in such remote areas as the islands of Southeast Asia. The social organisation and ritual customs of these modern farming communities are considered relevant when interpreting the archaeological evidence for early agricultural societies in Northern Europe.