Taphonomy and zooarchaeology of the Upper Palaeolithic cave of Dzudzuana,Republic of Georgia

We present the results of a detailed taphonomic and zooarchaeological study of the faunal remains from the Upper Palaeolithic layers of Dzudzuana Cave, Republic of Georgia. This study presents the first carefully analysed Upper Palaeolithic faunal assemblage from the southern Caucasus and thus serves as a significant point of reference for inter‐regional studies of Upper Palaeolithic subsistence in Eurasia. A series of intra‐site taphonomic comparisons are employed to reconstruct the depositional history of the bone assemblages within the different occupational phases at the site and to investigate subsistence, meat procurement and bone‐processing strategies. Caucasian tur (Capra caucasica), aurochs (Bos primigenius) and steppe bison (Bison priscus) were the major prey species throughout the Upper Palaeolithic. Their frequencies do not change significantly over time, and nor does bone preservation vary by layer. The assemblage is characterised by significant density‐mediated biases, caused by both human bone‐processing behaviours and in situ post‐burial bone attrition. Bone marrow extraction produced large numbers of unidentified bone fragments, many exhibiting green bone fractures. The density and size of bone assemblages and the extent of fragmentation indicate that Dzudzuana Cave was repeatedly occupied by Upper Palaeolithic foragers over many years. Skeletal part representation and butchery marks from all stages of carcass processing suggest that prey occasionally underwent field butchery. Intra‐site taphonomic comparisons highlight uniform patterns of cultural and economic behaviours related to food procurement and processing strategies. Copyright © 2007 John Wiley & Sons, Ltd.     

Taphonomy and zooarchaeology of a high-altitude Upper Pleistocene faunal sequence from Hovk-1 Cave,Armenia

The Caucasus is a key region for the study of hominid evolution and Neanderthal ecology. Taphonomic and zooarchaeological studies of sites from this region are few and only focused on sites at low-to-mid altitude zones with evidence of relatively intensive hominid occupation. This study focused on the taphonomic and zooarchaeological characteristics of a high-altitude site from the Upper Pleistocene – Hovk-1 Cave – looking at diachronic change in both natural and cultural processes which shaped the faunal assemblage. Results best fit a model in which the bones of most large mammals, mainly ungulates (wild goat, Capra aegagrus and red deer, Cervus elaphus) and cave bears (Ursus spelaeus) accumulated naturally through pitfalls, with minimal input from human or carnivore activity. This accumulation is characterized by a high frequency of complete ungulate and carnivore bones, a bear assemblage which is dominated by young-adults and a wild goat assemblage that includes juvenile and young-adult individuals. Our taphonomic reconstruction serves as a point of reference for comparative studies of palaeoenvironments and human subsistence patterns of Middle and Upper Palaeolithic sites in the Caucasus and broadens our perspective on hominid occupation and ecological adaptation in other high-altitude world regions.     

Final Paleolithic and Mesolithic Research in Reunified Germany

During the past decade research into the German Final Paleolithic and Mesolithic has experienced an important revival. One clear sign of this renewed interest in the periods are the annual meetings of the Arbeitsgruppe Mesolithikum (Mesolithic Working Group) which have taken place every spring since 1992. At these meetings, which take place at changing venues, topical themes of Final Paleolithic and Mesolithic interest are presented by informal lectures and it is also possible to study regional collections (artifacts, raw materials) at first hand. Numerous contributions were subsequently published together in one volume (Conard and Kind (1998) Aktuelle Forschungen zum Mesolithikum/Current Mesolithic Research, Mo Vince, Tübingen). The present paper intends to complement that collection of papers with a synthesis of developments and perspectives and to present recent research highlights in the German Final Paleolithic and Mesolithic, together with a comprehensive bibliography, to a wider international audience.     

Akhalkalaki: The Taphonomy of An Early Pleistocene Locality in the Republic of Georgia

As part of our investigations into the potential of the Republic of Georgia for providing information on early hominin occupation of Eurasia, we report here on Akhalkalaki, a large late Early Pleistocene locality located along the lower slopes of a Miocene andesitic cone. Originally excavated in the 1950s as a palaeontological site, it was re-opened in the 1990s and stone tools were found associated with the fauna, suggesting that it is also an archaeological occurrence. Excavations in the 1950s and 1990s uncovered thousands of bones of an early Galerian fauna, including the remains of new species of Hippopotamus,Equus , and Canis (Vekua, 1962, 1987) and dominated by the remains of Equus süssenbornensis. We present the stratigraphy of the site, which together with faunal correlations and reversed paleomagnetics indicates an age most likely in the late Matuyama Chron, probably between 980,000 and 780,000 years ago. Taphonomic analysis suggests that the fauna was deposited and buried over a short time period, and was heavily modified by carnivores, but we cannot demonstrate involvement by hominins. Based on evidence of abundant krotovina (animal burrows filled with sediment) and the lack of definitive evidence for hominin modification to the bones, the stone tools at the site may have been mixed in with the older fauna. The taphonomic characteristics of the Akhalkalaki bone assemblage are not readily explained with reference to assemblage formation processes developed with actualistic studies that have been mostly conducted in Africa, including carnivore dens, predator arenas, human hunting and scavenging, mass deaths, or attritional bone deposition. Because of extreme anthropogenic modification of the present environments, the temperate setting, and the presence of mainly extinct taxa, local models based on actualistic studies cannot approximate the mammalian ecology reflected in the Akhalkalaki bone assemblage. A few comparisons are made with preliminary taphonomic observations from Dmanisi, an Early Pleistocene Homo ergaster site not far away.     

Fossil fruit identification by SEM investigations, a tool for palaeoenvironmental reconstruction of Dmanisi site, Georgia

Archaeological excavations and multidisciplinary studies at the site of Dmanisi (Lesser Caucasus, Georgia) have provided an assemblage of lithic artefacts and numerous palaeontological and human remains in a volcano-sedimentary context dating to the early Lower Pleistocene. The sieving of archaeological sediments from Dmanisi also yielded mineralised botanical macro-remains that were analysed according to a particular protocol. The use of scanning electron microscopy (SEM) allowed the comparison between fossil fruit remains and modern specimens, leading to a precise identification of the earlier. Seed remains recognised by this method belonging to several taxa from the Boraginaceae family as well as to the genus Celtis L., of the Ulmaceae family. The ecological requirements of these taxa suggest the presence of xeric conditions, a result that is in accordance with data obtained by faunal and stratigraphic studies.     

Middle Holocene Taphonomy and Paleozoology at the Prairie-Forest Border,the Itasca Bison Site,MN

The Itasca Bison site near the prairie-forest border in central Minnesota is important to understanding the cultural and ecological role of bison in the upper Midwest during the middle Holocene. Recent reanalyses of this assemblage support the original interpretation of a taphonomically complex, fluvially modified site. Despite this taphonmic complexity, the role of humans in the butchering and possible accumulation of some of the bison fauna is also supported, although not as the only mechanism of accumulation. The bison assemblage is slightly biased toward males, and seasonality estimates are spread throughout the year. New direct accelerator mass spectrometry dates indicate a minimum of two highly mixed depositional events: one at 7970–7790 cal B.P. and an earlier event between 8520 and 8180 cal B.P.     

Excavations at the Bairaki Lower Paleolithic Site,Trans-Dniesteria,in 2011

Early Paleolithic deposits of the left-bank high seventh fluvial terrace of the Dniester were excavated at Bairaki, Trans-Dniestria. The assemblage is comprised of 28 artifacts including distinct flakes, cores, and Lower Paleolithic tools. Based on the 2011 geological and archaeological study, tools from the riverbed alluvium, paralleled by those of the Middle Oldowan, date to the earliest Lower Pleistocene. Bairaki, then, should be classified among the most ancient Paleolithic sites in Eastern Europe.     

Taphonomy and Paleoichnology of Olduvai Hominid 1 (OH1), Tanzania

This paper presents a case study of the Late Pleistocene skeletal remains of Olduvai Hominid 1, which were unearthed in Tanzania. Excavated in 1913, this fossil shows severe damage due to subterranean termites (Insecta: Isoptera). By combining methods to quantify and locate traces on the skeleton with the study of the different taphonomic processes that altered the deposit, we demonstrate a non‐arbitrary pattern of termite alteration of the skeletal remains. These results shed new light on the possibility to reconstruct the original position of a skeleton in a deposit and the potential to infer the space of decomposition and past funerary practices. Copyright © 2017 John Wiley & Sons, Ltd.     

Multiple origins of Bondi Cave and Ortvale Klde (NW Georgia) obsidians and human mobility in Transcaucasia during the Middle and Upper Palaeolithic

Using PIXE four types of elemental compositions were found among obsidian artefacts from the Bondi Cave and Ortvale Klde, Middle to Upper Palaeolithic sites in NW Georgia. One of those types corresponds to obsidians from the Chikiani source, whose compositions were determined with a very good agreement by PIXE and ICP-AES/MS. The composition of Chikiani obsidians is remarkably constant despite K–Ar and ³⁹Ar/⁴⁰Ar extrusion ages from ca 2.4 and 2.8 Ma. The compositions of two other groups of obsidian artefacts are similar to source materials from eastern Anatolia and Armenia, in particular Ikisdere, Sarikamis, Gutansar, and Hatis. Obsidian is only a minority component in the lithic assemblages at the Bondi Cave and Ortvale Klde. Both Neanderthal and Modern Human populations used obsidian in particular from Chikiani. Considering that the shortest walking distance to this nearest source is at minimum of about 180 km, and to other potential sources of more than 350 km it is suggested that this material reached these two sites mostly, if not exclusively, by a series of ‘down the line’ exchanges.     

Dental Remains from the Middle Paleolithic Layers of Altai Cave Sites*

Two Middle Paleolithic cave sites in the Altai – Okladnikov and Chagyrskaya – have yielded dental remains (mostly isolated teeth) of individuals of various ages. A newly discovered mandibular fragment with teeth from Chagyrskaya Cave reveals a Neanderthal trait combination: anterior fossa and epicristid (midtrigonid crest) on molars, metaconid and crest on premolars. The totality of dental traits support the conclusion previously drawn on the basis of postcranial characters: Altai Neanderthals appear to be intermediate between other Eurasian Neanderthals and anatomically modern humans.     

Carnivores and their prey in the Wezmeh Cave (Kermanshah,Iran): a Late Pleistocene refuge in the Zagros

Wezmeh Cave is located on the northeastern edge of the Islamabad plain, a high intermontane valley in the western‐central Zagros. In 1999 a disturbed but large faunal assemblage was recovered from this site. The abundant and extremely diverse faunal spectra present at Wezmeh Cave has highlighted the importance of this assemblage. Carnivore remains constitute the bulk of the assemblage; red fox (Vulpes vulpes) has the highest number of identified specimens followed by spotted hyena (Crocuta crocuta), brown bear (Ursus arctos), wolf (Canis lupus), felids (lion, leopard, lynx/caracal and wildcat), mustelids (badger, polecat, marten) and viverrids (mongoose). Artiodactyls (bovid, cervid, suid), equids, rhinoceros (Dicerorhinus sp.) and small animals (Cape hare, porcupine, tortoise, snake, birds) are also present. According to U‐series dating, the site was occupied from around 70 ka BP through to sub‐recent periods by carnivores. Amongst this rich assemblage, a human fossil tooth was also found and dated by non‐invasive spectrometry gamma dating to 20–25 ka BP. A preliminary zooarchaeological and taphonomic study shows that Wezmeh Cave was used by multiple carnivore species, a unique phenomenon in the Zagros Mountains in particular and southwest Asia in general. Copyright © 2008 John Wiley & Sons, Ltd.     

Taphonomic and Methodological Perspectives of Leporid Hunting During the Upper Paleolithic of the Western Mediterranean Basin

Leporid (rabbit and hare) bones have been shown to yield important information about subsistence practices, mobility patterns, and demographic trends during the Paleolithic of the western and eastern Mediterranean regions. Studies of Spanish Paleolithic caves rich in rabbit bones suggest that residential mobility patterns influence the degree of leporid hunting through time. Studies of Paleolithic sites in the eastern Mediterranean suggest that leporids were hunted in large numbers only after population sizes and densities reached certain thresholds. This paper reviews and critiques these studies based on current taphonomic and ecologic information about leporids. Leporid hunting during the Upper Paleolithic of central Portugal is then discussed and compared to these existing models. These latter data suggest that rabbit hunting in central Portugal does not conform to any existing model, suggesting that local factors of leporid density and environmental conditions likely influenced the nature and timing of small game acquisition during the Upper Paleolithic.     

The Neanderthals of Okladnikov Cave Altai: Environment and Diet Based on Isotopic Analysis*

The discovery of a Mousteroid industry in the northwestern Altai represents a major finding in Eurasian Paleolithic studies. Isotopic analyses of bone collagen of late Western European Neanderthals have repeatedly indicated that they were extremely carnivorous. However, no corresponding information was previously available for the Neanderthals of the Altai. With this in mind, an isotopic study was conducted to assess the diet of the Middle Paleolithic inhabitants of Okladnikov Cave. Collagen was extracted from seven well preserved bone samples of adults and children and 13С/12С and 15N/14N ratios were calculated. The δ13С content suggests that all individuals were members of a local trophic chain based on plants of the moderate climatic zone. However, the δ15N content demonstrates that their trophic level was high. These findings suggest that the lifestyle of Neanderthals in the northwestern Altai was similar to that of the late European Neanderthals. Both evidently subsisted by hunting large herbivores.     

The Taphonomy of the Final Middle Stone Age Fauna from Sibudu Cave,South Africa

This study applies a taphonomic analysis to the final Middle Stone Age faunal assemblage from Sibudu Cave, South Africa, by assessing bone surface modifications, breakage patterns and skeletal element abundances. Cut marks, percussion marks, severe fragmentation and the high frequency of burned bone combine to demonstrate that human behaviour was the principal agent in the assemblage's formation. These results are consistent with previous research on earlier occupations of Sibudu during the Middle Stone Age. Moreover, this assemblage is proposed to reflect regular site maintenance and cleaning. This conclusion is consistent with previous research that demonstrates systematic site maintenance during the Middle Stone Age at Sibudu and emphasises this behaviour as being a consistent activity for Middle Stone Age foragers. Copyright © 2013 John Wiley & Sons, Ltd.     

Modern humans in Sarawak,Malaysian Borneo,during Oxygen Isotope Stage 3: palaeoenvironmental evidence from the Great Cave of Niah

During recent reinvestigations in the Great Cave of Niah in Borneo, the ‘Hell Trench’ sedimentary sequence seen by earlier excavators was re-exposed. Early excavations here yielded the earliest anatomically-modern human remains in island Southeast Asia. Calibrated radiocarbon dates, pollen, algal microfossils, palynofacies, granulometry and geochemistry of the ‘Hell Trench’ sequence provide information about environmental and vegetational changes, elements of geomorphic history and information about human activity. The ‘Hell’ sediments were laid down episodically in an ephemeral stream or pool. The pollen suggests cyclically changing vegetation with forest habitats alternating with more open environments; indicating that phases with both temperatures and precipitation reduced compared with the present. These events can be correlated with global climate change sequences to produce a provisional dating framework. During some forest phases, high counts of Justicia, a plant which today colonises recently burnt forest areas, point to fire in the landscape. This may be evidence for biomass burning by humans, presumably to maintain forest-edge habitats. There is evidence from palynofacies for fire on the cave floor in the ‘Hell’ area. Since the area sampled is beyond the limit of plant growth, this is evidence for human activity. The first such evidence is during an episode with significant grassland indicators, suggesting that people may have reached the site during a climatic phase characterised by relatively open habitats ∼50 ka. Thereafter, people were able to maintain a relatively consistent presence at Niah. The human use of the ‘Hell’ area seems to have intensified through time, probably because changes in the local hydrological regime made the area dryer and more suitable for human use.     

Herpetofaunal Remains from Khonkho Wankane,an Urban and Ceremonial Centre in the Southern Lake Titicaca Altiplano: Unique Behavioural Correlates to Osseous Taphonomy

This research reports the herpetological (reptile and amphibian) remains from Khonkho Wankane, an urban and ceremonial centre in the southern Lake Titicaca region of the Bolivian Altiplano, which fluoresced during the Late Formative period. The total n = 1710, including a minimum of six taxa and representing a significant portion of the relatively depauperate modern herpetofauna recorded for the Lake Titicaca area. A lizard of the genus Liolaemus (Tropiduridae) was identified. Amphibians are represented by Andean toad Rhinella spinulosa (Bufonidae) and by at least four taxa of frogs: [cf. Gastrotheca marsupiata (Hylidae) and Pleurodema marmorata or P. cinerea and Telmatobius spp. (Leptodactylidae)]. The latter genus does not include members of the giant variants (T. culeus) that dwell in Lake Titicaca. The question of how these remains were incorporated into the site deposits is examined taphonomically. No indicators of human or small carnivore predation were detected. Significant spatial clustering of amphibian remains was detected, with two clusters contributing n = 1562 or 92.4% of the amphibian total (n = 1690). These concentrations also are characterised by a depressed ratio of cranial versus postcranial elements, which cannot be explained by recovery methods. These concentrations of herpetofaunal remains likely represent in‐burrow deaths of species sheltering for thermoregulatory benefits away from this harsh Altiplano environment, with the lowered recovery of cranial elements hypothesised to have resulted from a lack of ossification. The bulk of herpetofaunal remains recovered therefore likely were recent intrusions into the site deposits and do not represent the residue of human exploitation during the occupation of Khonkho Wankane. Copyright © 2011 John Wiley & Sons, Ltd.     

Late Middle and Early Upper Paleolithic Evidence from the East European Plain and Caucasus: A New Look at Variability,Interactions, and Transitions

This paper presents analyses of Late Middle Paleolithic (LMP) and Early Upper Paleolithic (EUP) material from the East European Plain and Caucasus. Late Middle Paleolithic industries show a highly variable pattern, although they are formally ascribed to a limited number of technocomplexes. Many of the LMP industries, especially in the Crimea, survived to the time of the transition to the Upper Paleolithic, but data suggesting a local origin of EUP are extremely rare. The transition is generally dated between 32/30,000 and 26/24,000 years, while the most crucial changes coincide with the Stillfried B interstadial. Aurignacian (two variants), Gravettian, and Transitional industries are recognized in the EUP. The presence of Middle Paleolithic traits in the Aurignacian may indicate acculturation, while the Transitional industries might reflect either acculturation or independent local development of new technologies, raising the possibility of local transformation of some Middle Paleolithic into non-Aurignacian EUP industries.     

Exchange patterns,boundary formation,and sociopolitical change in Late Bronze Age Southern Caucasia: preliminary results from a pottery provenance study in northwestern Armenia

In the South Caucasus—roughly the territory of today's Armenia, Georgia, and Azerbaijan—the transition from the Middle Bronze Age (MBA) to the Late Bronze Age (LBA) is equated with fundamental shifts in settlement patterns, subsistence economy, and political strategies. During the mid-2nd millennium BC, nomadic pastoral societies that had dominated the region began to settle down and construct stone fortresses along the foothills of the Lesser Caucasus; these fortifications largely replaced the expansive and often opulently adorned kurgan burials as the most prominent expression of political dominance on the landscape. After a decade of intensive archaeological study at various fortifications, very little remains known about the political and economic relationships among fortresses on a regional scale that might improve our understanding of the roots of these sociopolitical transformations. In this paper, we highlight the results of a recent neutron activation analysis (NAA) of ceramics from elite and non-elite contexts at a selection of LBA fortresses on the Tsaghkahovit Plain in northwestern Armenia, and offer some preliminary interpretations about political and economic organization and boundary formation. Most strikingly, the NAA data suggest that the fortresses on the Tsaghkahovit Plain appear to have isolated themselves economically from surrounding valleys, perhaps in an attempt to forge boundaries and legitimating ideologies attendant to new political formations that were quite distinct from their nomadic predecessors in the MBA.     

Taphonomy,life history,and human exploitation of Rhinoceros sinensis at the Middle Pleistocene site of Panxian Dadong,Guizhou, China

Rhinoceros remains are commonly found in Chinese Pleistocene archaeological localities. This study examines the characteristics of the Rhinoceros sinensis sample from Panxian Dadong, a karst cave in the mountains of western Guizhou province, with a mammalian fauna in association with stone artefacts and human remains from the late Middle Pleistocene (MIS 6‐8). The distribution of skeletal elements shows a predominance of foot (metapodial and phalanges) and lower limb (carpals and tarsals) bones, while the dental age‐at‐death profile, constructed using dental eruption and tooth wear data, is characterised by a high frequency of prime age adult teeth. There is little taphonomic evidence for the involvement of non‐human carnivores or natural agencies in the formation of the faunal assemblage. Instead, it appears that human activities were responsible for the unexpected prevalence of prime age adults. Copyright © 2008 John Wiley & Sons, Ltd.