The taphonomy and palaeoenvironmental implications of the small mammals from Karain Cave,Turkey

The small mammal accumulations in the Pleistocene deposits of Karain Cave are investigated to identify the predators and possible biases in the fossil assemblages. Seven small mammal assemblages are studied in chronological order from two chambers of the cave, the main chamber E and the side chamber B. The lowermost level within the whole sequence is Proto-Charentien, which corresponds to an early stage of the Middle Palaeolithic. The main part of the material belongs to Middle Palaeolithic layers. The most important aspect of the fossil record in the cave is the human occupation without any interruption through the Pleistocene to Holocene. The small mammal fossil evidence in the cave denotes the presence of opportunistic predators throughout the sequence with one exception, and these produce balanced samples of small mammal faunas in the habitat. The lack of bias in the small mammal faunas allow the interpretation of local environments, showing that partial steppe and arid conditions existed during deposition of the lowermost levels of the Middle Palaeolithic in Karain Cave and that these shifted into more temperate and wooded habitat in the upper levels. Evidence from the side chamber indicates some differences, with a more open grassy environment. In the Mediterranean part of Anatolia the temperate and moist conditions in the Middle Palaeolithic were superseded by more arid conditions in the Upper Palaeolithic, followed by a decrease in steppe conditions during the Epi-Palaeolithic period.     

The British Late Middle Palaeolithic: An Interpretative Synthesis of Neanderthal Occupation at the Northwestern Edge of the Pleistocene World

The British Middle Palaeolithic is divided into two discrete periods of occupation: the Early Middle Palaeolithic (MIS 9–7, ~330–180 ka BP) and the Late Middle Palaeolithic (MIS 3, ~59–36 ka BP), separated by a long hiatus. Owing to the relative poverty of the record and historical difficulties in dating and correlating archaeological sites, the British Late Middle Palaeolithic has, until recently, received scant attention, and has largely been regarded as the poor man of Europe, especially by British archaeologists. Indeed, there has been more discussion of the absence of humans from Britain than of what they did when they were present. We aim here to redress that situation. Following from recent considerations of the Early Middle Palaeolithic (White et al. in J. Quat. Sci. 21:525–542, 2006; Scott, Becoming Neanderthal, Oxbow, Oxford, 2010), we offer an interpretative synthesis of the British Late Middle Palaeolithic, situating ‘British’ Neanderthals in their chronological, environmental and landscape contexts. We discuss the character of the British record, and offer an account of Neanderthal behaviour, settlement systems and technological practices at the northwestern edge of their known Upper Pleistocene range. We also examine the relationship of the enigmatic Early Upper Palaeolithic leafpoint assemblages to Neanderthals.     

The black layer of Middle Palaeolithic combustion structures. Interpretation and archaeostratigraphic implications

Certain aspects of the formation processes of simple, flat archaeological combustion structures such as those present in the Middle Palaeolithic record remain unexplained. Such kind of combustion structures are commonly affected by postdepositional agents and often, their only distinct, well preserved component is a thin black lens on the ground. Hence, understanding the nature of this black lens is essential towards archaeological interpretation. From an interdisciplinary microstratigraphic approach, we present a case study in which for an entire experimental series of flat combustion structures the black layer represents the fire-altered topsoil on which the fire was made. Parallel analysis of archaeological Middle Palaeolithic combustion structures from the site of El Salt (Alicante, Spain) reveal similar patterns, leading to significant implications for archaeological interpretation. In the light of these results, special attention must be paid to the formation processes of flat Middle Palaeolithic combustion features, as black layers and the material contained in them are not necessarily linked with combustion but with preceding activities or events. In such cases, black layers represent intact remnants of occupation surfaces, concealing significant behavioural and palaeoenvironmental information relevant to the reconstruction of Middle Palaeolithic societies.     

Using lithic assemblage formation to approach Middle Palaeolithic settlement dynamics: El Salt Stratigraphic Unit X (Alicante,Spain)

The superposition of anthropogenic remains usually results in archaeological palimpsests, hindering an accurate characterization of the behavior underlying such remains. Aimed at facing this methodological constraint in the study of Palaeolithic contexts, we present a contextual approach to Neanderthal settlement dynamics based on an understanding of lithic assemblage formation processes. We focused on a set of lithic artifacts from the Middle Palaeolithic site of El Salt. With the goal of isolating temporally constrained units of behavioral analysis, we analyzed the spatial distribution of lithic, faunal and combustion remains. Then, the lithic records from each of the resulting units were analyzed from a genetic and technical perspective. This goal was achieved by identifying lithic Raw Material Units (RMU) through observations on reduction processes, refits and their spatial distribution. Our results suggest that El Salt SU X lithic remains represent several diachronic human occupation episodes and different site functions. This study illustrates the benefit of investigating lithic assemblage formation processes to alleviate the palimpsest problem in Palaeolithic contexts.     

Trampling experiments at Cova Gran de Santa Linya,Pre-Pyrenees,Spain: their relevance for archaeological fabrics of the Upper–Middle Paleolithic assemblages

The study of fabrics, that is, the analysis of the orientation and slope of archaeological and sedimentary materials associated with the Middle Palaeolithic/Upper Palaeolithic (MP/UP) transition at Cova Gran shows substantial differences. Archaeological assemblages are characterised by greater isotropy in the fabrics than the sedimentary levels within which they are located, indicating that these differences may be generated by anthropic processes. One of the anthropogenic processes associated with horizontal and vertical displacement of archaeological artefacts is trampling and circulation caused by later occupations. In order to evaluate the effect of movement on materials, we undertook experiments simulating geological and archaeological conditions at Cova Gran. The results show that human trampling does not cause major isotropy in fabrics, but arranges archaeological assemblages towards planar or linear materials according to surface geometry. We were not able to replicate the fabric pattern of materials from the archaeological levels of Cova Gran, suggesting that they must be associated with the activities of human occupation at each level.     

Did Modern Human Dispersal Take a Coastal Route into India? New Evidence from Palaeolithic Surveys of Kachchh,Gujarat

The Indian Ocean coastline is argued to have been a critical route of modern human dispersal from Africa, introducing Late Palaeolithic industries into South Asia, but a dearth of evidence has prevented a direct evaluation of this. Kachchh (Gujarat, India), located immediately east of the Indus Delta, is an important setting to appraise the Palaeolithic occupation of the western Indian coastline. Targeted survey of Late Pleistocene sediments there found widespread evidence for occupation of Kachchh during the Late Pleistocene: Middle Palaeolithic and possibly Late Acheulean lithic artifacts. The conspicuous absence of Late Palaeolithic industries in these Late Pleistocene deposits, with their presence only noted in Holocene contexts, does not support the model of modern human expansions into India with these industries via the coastal route. We evaluate these results in the context of current debates regarding Late Pleistocene hominin demography, adaptation, and expansions.     

Points and convergent tools in the European Early Middle Palaeolithic site of Payre (SE,France)

The lithic assemblage from the Middle Palaeolithic site of Payre (Rhone Valley, France) contains a large number of convergent tools and pointed tools of various shapes, sizes and retouch types. These were excavated from several archaeological units, dating from marine isotopic stages 8–5, which also yielded human skeletal remains. Consideration of this large tool kit has led to an improved analysis of Middle Palaeolithic tools with two retouched convergent edges. The 350 tools were not described within the classical typological framework, but, rather, from a lithic technological perspective in relation to a discoid debitage. In addition, an initial macroscopic use-wear analysis aided in establishing whether they were used according to their technical and/or morphological features. The Middle Palaeolithic convergent tools from Payre are shown to be quite diversified, and the question of the significance of the retouch and the definition of the various types is addressed. Initial functional results indicate that a clear relationship between shape and function cannot be easily established, and that these tools were used as hand tools. This study contributes to the debate on the use of stone tip spears in the Early European Middle Palaeolithic.     

The Palaeolithic of the Middle Son valley, north-central India: Changes in hominin lithic technology and behaviour during the Upper Pleistocene

The Middle Son valley in north-central India preserves extensive Quaternary alluvial deposits. A long history of archaeological and geological research in the valley has resulted in the discovery of lithic assemblages ranging from Lower Palaeolithic to microlithic, a rich corpus of fossilised faunal remains, and ash deposits from the 74,000 year-old Toba supereruption. This paper reviews the chronology and stratigraphy of the valley’s Quaternary sediments, and presents a model that hypothesizes the temporal sequence of important lithic assemblages from excavated and surface contexts. Artefacts in these assemblages are analysed and changes in lithic technology through time are described; this evidence is used to propose shifts in hominin behaviour and demographic structure in this region during the Upper Pleistocene. Recognising gaps in our understanding of the Middle Son record, future avenues of research are recommended that will build upon previous research and address questions of palaeoanthropological significance. The Middle Son valley preserves a long and rich record of hominin occupation from all periods of the Palaeolithic that is rarely paralleled by other sites in India.     

Developmental Stress in Immature Hominines from Late Pleistocene Eurasia: Evidence from Enamel Hypoplasia

The differences between Middle and Upper Palaeolithic cultures from Europe have been likened to the contrast between generalized and complex hunter–gatherers. A test of this model is undertaken by comparing the types, amount, severity and timing of episodes of enamel hypoplasia between the two periods among specimens from western Europe and the circum-Mediterranean area. The earlier sample consists of 59 dentally immature individuals with 128 primary and 154 permanent teeth while the later sample consists of 47 dentally immature individuals with 162 primary and 125 permanent teeth. It was predicted that the Upper Palaeolithic would show more variation in the attributes noted above as a consequence of increased social differentiation. While the prevalence (50% of individuals) and severity of enamel hypoplasia do not differ between the samples, the Upper Palaeolithic component is significantly more variable in the types and timing of enamel defects. Localized hypoplasia of the primary canine is absent from the Middle Palaeolithic but very common among Upper Palaeolithic children. Earlier work has linked this enamel defect to craniofacial osteopenia due to low bioavailability of vitamin A. Linear enamel hypoplasia peaks at age 3·5 years in the Middle Palaeolithic sample but is as common at age 2·0 as 5·0 years in the Upper Palaeolithic sample. The age distribution differs significantly. Peak age at stress, combining all types of enamel hypoplasia, is 3·5 years in the Middle Palaeolithic and 0·5 years in the Upper Palaeolithic. The major contrast between the Middle and Upper Palaeolithic in terms of enamel hypoplasia is the advent of stressful episodes in early infancy in the Upper Palaeolithic. This is attributed to the low bioavailability of vitamin A due to the synergistic effects of malnutrition and infection exacerbated by a net increase in population density among socially competitive family lineages.     

Reconnaissance survey for Palaeolithic sites in the Debed River Valley,northern Armenia

Abstract

The southern Caucasus is a critical region for those interested in Palaeolithic research because of its varied topography and location at the crossroads of Europe, Africa, and Asia. Modern Armenia sits at the heart of this area, but has until now played a small role in broader debates, largely because of its paucity of well-excavated and well-dated sites. To improve this situation, a survey was conducted for Palaeolithic sites along the valley of the Debed River (Lori Depression, northeastern Armenia). Twenty-three open-air sites, spanning the Lower through the Upper Palaeolithic periods, were identified. Most of the lithic material is of Middle Palaeolithic manufacture. Upper Palaeolithic material is also well represented, but only a handful of Lower Palaeolithic artifacts have been identified. Test excavations at several sites suggest that they preserve in situ deposits that may help us to understand the role of the southern Caucasus in the Palaeolithic occupation of Eurasia.     

The Neronian and the historical structure of cultural shifts from Middle to Upper Palaeolithic in Mediterranean France

The end of the Middle Palaeolithic period can be considered a key moment in the history of humanity, characterized by profound changes in traditional Palaeolithic societies. Whatever the reasons for this historical change, it seems well established that it took place within a particularly dynamic phase of these societies, when Upper Palaeolithic technical processes were already emerging. The analysis of recent Mousterian industries in Mediterranean France highlights certain points of inflection, which presage new ways of life. The analysis of key sites demonstrates important transformations within the last Neanderthal societies, with a gradual rearticulation of Mousterian technical systems around the production of blades and points and with profound changes in the relationship of the craftsman with his tools. Even if they are typical of the late Mousterian more generally, these technically distinctive industries have such distinct chronological, territorial and cultural characteristics that they can be referred to by a specific name, the “Néronien” (Neronian). Stratigraphically speaking, this group of assemblages – at the crossroads of the Middle and Upper Palaeolithic – appears in some important regional sequences just before the appearance of the first “true” Upper Palaeolithic industries (the Protoaurignacian). However, this Neronian doesn't represent the end of the Middle Palaeolithic period in the Rhône valley. The final Moustérien of this region is actually marked by deep breakdowns of the traditional balances in the human groups. This second point of inflection could characterize a true destabilization of Mousterian societies. There is an interesting parallelism between the historical paths of the Castelperronian in the arch of the Massif Central and the Neronian in the Rhône valley. It also highlights the mosaic nature of the profound cultural changes that occurred at the end of the Middle Palaeolithic.     

Taphonomic resolution and hominin subsistence behaviour in the Lower Palaeolithic: differing data scales and interpretive frameworks at Boxgrove and Swanscombe (UK)

The identification of human butchery-signatures on fauna from Lower Palaeolithic sites is well documented and readily identifiable. Such bone surface modifications have the potential to provide not only information about past hominin meat-procurement behaviour but address the wider issue of competition for resources with other carnivore species. To understand and discuss these broader issues both hominin and natural bone surface modifications must be understood and contextualised within a site-specific spatial and temporal framework. This paper presents new results from faunal analysis at two key British Lower Palaeolithic localities: Boxgrove and Swanscombe. It illustrates that different depositional environments and excavation histories have resulted in different scales and resolutions of available data and hence in varying interpretive potentials. At Swanscombe the archaeological record has been disturbed by both fluvial activity and excavation history providing a coarser-grained record of anthropogenic behaviour than previously acknowledged. Conversely, at Boxgrove, a finer-grained, higher resolution record of human behaviours has been preserved; this, combined with both an extensive and intensive excavation strategy, has allowed for a broader discussion of hominin landscape use, resource competition and meat-procurement behaviour. This paper highlights that assessing the specific depositional environment at each site is crucial to understanding Palaeolithic faunal assemblage formation and, consequently, the available data-resolution and behavioural interpretation.     

The Muddle in the Middle Pleistocene: The Lower–Middle Paleolithic Transition from the Levantine Perspective

The terms Lower Palaeolithic and Middle Palaeolithic represent research constructs within which cultural evolution and prehistoric hominin behaviours can be studied, with the transition usually understood as marking a watershed in our evolution: an adaptation with a million-year record of success that gives way to something new. The interpretation of the Lower Palaeolithic Acheulian technocomplex is usually understood as a period of cultural stasis that extends over much of Africa and Eurasia, principally associated with Homo erectus. Those innovations that can be observed occur widely separated from one another in space and time. Yet a closer and more detailed examination of the Middle Pleistocene records from East Africa, southern Africa, Europe and the Levant reveals significant variation in cultural repertoires. A kind of paradox emerges, in which an Old World Lower Palaeolithic, apparently lacking an overall dynamic of distinctive and directed change in terms of cumulative variation over time, nevertheless culminates in a transition which sees the universal appearance of the Middle Palaeolithic. The two main hypotheses that have been advanced to explain the global transition, which happens essentially synchronously, appear mutually exclusive and contradictory. One view is that altered climatic-environmental constraints enabled and encouraged an ‘Out-of-Africa’ dispersal (or dispersals) of a new type of genus Homo. This cultural replacement model has been challenged more recently by the alternative hypothesis of accumulating but unrelated and temporally non-linked regional, and in fact potentially autochthonous, processes. The Levant, by virtue of its position bridging Africa and Eurasia (thus being the region into which any out-of-Africa groups would have had first to disperse into), must be seen as a critical region for assessing the relative merits of these competing hypotheses. This paper deals with the Lower–Middle Paleolithic boundary in the Levant within a long temporal perspective. The Middle Pleistocene record in the Levant enables us to examine the amplitude of variation within each techno-complex, as well as to question whether there are diachronic changes in the amplitude of techno-typological variations as well as changes in the manner by which they appear in the record. The results carry significant implications for understandings of demographic and societal processes during the Lower–Middle Paleolithic transition in the Levant.     

The Definition of Spatial Units in Middle Palaeolithic Sites: The Hearth-Related Assemblages

This paper deals with hearth-related accumulations in Middle Palaeolithic sites. Spatial behaviour is a key to understanding the cultural capabilities of ancient hominids. Ethnoarchaeological and archaeological research has shown that hearth-related assemblages are a basic feature in the spatial behaviour of modern and prehistoric hunter–gatherers. In this paper, we propose a methodology for analysing archaeological accumulations and study a series of hearth-related assemblages from the Abric Romanı́ Middle Palaeolithic site (Capellades, Spain). We also analyse the degree of contemporaneity between the different activity episodes documented at each archaeological accumulation. Our results suggest that the characteristics of domestic areas in this site are not substantially different from those observed in ethnoarchaeological contexts and Upper Palaeolithic sites.     

Palaeolithic Human Subsistence in Mount Carmel (Israel). A Taphonomic Assessment of Middle and Early Upper Palaeolithic Faunal Remains from Tabun,Skhul and el‐Wad

Detailed archaeozoological studies of Middle and Early Upper Palaeolithic (250–40 kya) faunal assemblages from Tabun, Skhul and el‐Wad caves, located in Mount Carmel (Israel), are presented here, expanding on the previous palaeontological analyses carried out by Garrod and Bate in the 1930´s and by Garrard five decades later. Despite the well‐known excavation bias of the older excavations, this study makes greater use of modern taphonomic techniques and statistical procedures, providing new palaeoeconomic insights into the subsistence behaviour of modern humans, Neanderthals and their predecessors in the Levant. The results show that anatomically modern humans (AMH) were more efficient at hunting and foraging in terms of human mobility as well as an incipient prey specialisation. In addition, different uses of the Tabun cave through time have been identified. Tabun B, initially considered as a palaeontological accumulation, was also occasionally accessed by Neanderthals in order to exploit naturally trapped ungulates, and there were sporadic and logistic occupations in Tabun D, one of the oldest Middle Palaeolithic assemblages in the area. Contrastingly, Tabun C showed a comparatively more residential use, which coincided with the new mobility capabilities of AMH. Whereas the intensity of human occupation at el‐Wad varied through time, levels G and F showed very brief occupations in comparison with levels D and E, which showed abundant evidence of human activity. Copyright © 2011 John Wiley & Sons, Ltd.     

Radiocarbon dates for the late Middle Palaeolithic at Pech de l'Azé IV,France

This paper reports a series of radiocarbon dates on bone samples coming from the Mousterian of Acheulian Tradition (MTA) Layer 3 at the top of the Pech de l'Azé IV (Pech IV, France) Middle Palaeolithic sequence. All of these samples showed evidence of human impact, and they were prepared using current pre-treatment techniques to remove or identify contamination. The results indicate that the MTA at Pech IV started prior to the current limit of radiocarbon calibration (circa 50 ka BP) and ended by 45 ka cal. BP. These dates are supported by additional TL and ESR dates from the sequence, confirm previously suggested correlations between Pech I and IV, and generally fit within the known age range for the MTA. The oldest dates reported here may also lend support to still older TL dates for the MTA that taken together suggest that the MTA extended from late MIS 4 or early OIS 3 through to the end of the Middle Palaeolithic in southwest France.     

On Neanderthal subsistence strategies and land use: A regional focus on the Rhone Valley area in southeastern France

This study of Middle Palaeolithic assemblages from the Rhone Valley, in the South-East of France, increases our understanding of Neanderthal subsistence strategies and modes of territorial organisation by comparing a wide corpus of human occupations in limited chronological and geographical frameworks. The Neanderthal occupation modes may be examined using sites located in a reduced area, linking medium-altitude territories (Massif Central and the Alpine foothills) to the Rhone corridor. Through the combined analysis of the occupation levels of ten sites, all dated to between Marine Isotopic Stage (MIS) 7 and the beginning of MIS 3, we identify three types of occupation durations: (1) type 1 as long-term residential camps, (2) type 2 as short-term regular hunting camps and (3) type 3 as brief stopover camps. We discuss this variability of habitat types according to various parameters: site age, technical behaviour, environmental conditions, and site localisation and occupation seasons. The aim is to discern the underlying motivations behind Neanderthal group mobility. One of the main features of the Rhone Valley area is the great homogeneity of behaviours reflected within the sequences. This homogeneity, linked to the variability of the site occupations, supports the hypothesis of Neanderthal groups anticipating their land use requirements, and furthermore suggests that another type of circulating model was used in this area.     

New Middle Palaeolithic sites from the Mani Peninsula,Southern Greece

We here report the first results from a systematic research project in Mani (Southern Greece), which includes survey and test excavations. Forty-six caves, rockshelters and open-air sites in lowland settings were surveyed. Geomorphological data were collected in order to assess how geological processes affect the preservation of sites and bias site distribution patterns. Artifacts manufactured from non-local rock indicate potential raw material transfers and suggest links among the different regions of Mani, related to mobility patterns. Our research in the Mani has nearly doubled the number of known Middle Palaeolithic sites from the region and confirmed that the peninsula has the strongest ‘Neanderthal signal’ identified to date in Greece. Almost all sites are located at coastal areas. Despite the influence of Pleistocene landscape dynamics, this distribution emerges as a persistent pattern, perhaps indicating a preference for coastal locations. The Neanderthal occupation of Mani can illuminate important aspects of Middle Palaeolithic adaptation in one of the southernmost coastal regions of Europe.     

Seasonality and intra-site variation of Neanderthal occupations in the Middle Palaeolithic locality of Payre (Ardèche,France) using dental wear analyses

The reconstruction of Palaeolithic subsistence and settlement systems at the Middle Palaeolithic site of Payre (France) is undertaken through the application of dental wear analyses combined to zooarchaeological, technological, and ecological indicators. Three archaeological levels were investigated. Level D, dated to MIS 5, could correspond to an occupation during the cold season. In levels G and F, dated to MIS 8-7, animals were probably hunted during a warmer season. According to dental wear analyses, it is likely that level F has actually recorded a succession of short occupations, contrary to what is observed in the levels G and D. Those differences in the duration of site occupation are discussed in the light of previous sedimentological, zooarchaeological, and technological studies.     

Mixed occupation during the Middle Palaeolithic: Case study of a small pit-cave-site of Les Pêcheurs (Ardèche,south-eastern France)

The Pêcheurs cave is a unique example of a Middle Palaeolithic site with three kinds of accumulations: (1) ibex that died in a natural trap, (2) carnivores that died within the cavity, and, (3) a series of short-term occupations by humans who left a few artifacts and a hearth area. Biological patterns of ibex remains (skeletal parts, age) show a homogeneous structure, related to natural death inside the cave. The Chassezac valley is narrow and sinuous, bordered by steep cliffs occupied by well-adapted hoofed-species (Caprinae). Moreover, Les Pêcheurs is a shallow cave, pit-like, and in its deepest part (Sector 4) provided both man and animals with shelter. The presence of a fire place (in the middle of the sequence of Sector 4) firmly indicates the presence of an in situ occupation by a small group of hominids. According to the stratigraphical patterns and the analysis of the lithic assemblages, artifacts do not seem to have been introduced into sediments. The lithic assemblages (technically homogeneous) indicate that small mobile human groups inhabited a cave that offered, by virtue of its morphology, a natural shelter against the cold winds blowing in the Chassezac valley and the plateaus of the south-eastern borders of the Massif Central Mountains. The exploitation of biotopes such as this rocky area constitutes a specific case of human subsistence behavior and settlement strategy. The deepest layer is characterized by a lithic assemblage mostly made of local raw material (quartz) implying a low investment in lithic production. Few flakes made from non-local flints attest to the mobility of the human occupants who moved across these areas and perhaps found in the valleys, short-term refuge.