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1.
This study documents and interprets adaptive postcranial morphology among prehistoric Jomon period foragers from Hokkaido, Japan (HKJ). The Hokkaido climate is differentiated from other Japanese islands by freezing winters with sea‐ice accumulation in the northern regions. Increased brachial and crural indices are, however, observed among HKJ foragers, while body mass (BM) has not yet been estimated for these groups. Based on previous observations and paleoclimatic reconstructions, it was predicted that increased BM and increased distal relative to proximal limb lengths would typify HKJ foragers. Similar BM was observed between HKJ foragers and groups from colder environments. Intralimb indices do, however, suggest similarity between HKJ foragers and groups from high‐latitude, warm environments. It is likely that HKJ foragers retained cold‐derived BM in association with Pleistocene migrations to Hokkaido via Northeast Asia. That is, enlarged BM among HKJ foragers is associated with long‐term evolution in a colder environment. Relatively elongated distal limbs may represent morphological response to a slightly warmer environment. Following migration to Japan from a colder environment, elongation of distal limb segments resulted in elevated brachial and crural indices. Relatively elongated distal appendages may also reflect positive nutritional status as HKJ people experienced lesser rates of systemic stress than other Jomon groups. It is also possible that elongated distal relative to proximal limbs are associated with neutral mutation and genetic drift. This interpretation suggests a neutral mutation associated with relative limb length in some HKJ ancestor with subsequent spread of this allele through isolation and drift. Ontogenetic and temporal studies of intralimb indices among Jomon people are necessary to further evaluate these interpretations. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

2.
The long bones of the human upper limb usually show lateral asymmetries of length. This pattern can be attributed either to the mechanical consequences of handedness bias or to genetic or hormonal factors acting directly on longitudinal bone growth. Length data was obtained from the long bones of the upper limbs of a large skeletal assemblage from Wharram Percy, Yorkshire (England), predominantly deriving from the 11th-16th centuries A.D. The Wharram Percy adult skeletons had a population distribution of lateral asymmetries of length in the humerus and in the humerus-plus-radius (a proxy arm length index) which closely parallels the pattern of behavioural handedness found in modern populations. This pattern was developing in the skeletons from the infant and juvenile age ranges, but was absent in the neonates (of whom 12 out of 14 had longer left humeri). We argue that this supports the environmental hypothesis that the ontogeny of long bone length asymmetry is consequent to the earlier development of lateral bias in mechanical loading of the upper limbs.  相似文献   

3.
This paper attempts to define the relation between (1) human skeletal measurements and (2) height, weight and physique indices. Admitted to the study were 686 men and women from five ethnic groups (White, Inuit, Gurkha, Bantu and San). In addition to height and weight, subjects had defined cranial and postcranial measurements taken using methods derived from clinical examination techniques. Measurements were checked for reliability. Skinfold callipers were used to allow for the thickness of soft‐tissues overlying bone widths. Adjustment factors were derived from radiological gold standards. Radiological measurements, adjusted for magnification, of the lumbar vertebrae from 42 subjects were also done. A correlation matrix was constructed for the complete study population and all variables to allow a preliminary assessment of useful predictive independent variables. Discriminant scores for classification by gender were calculated. Regression computations, using least square regression, were calculated for six defined reference populations. The regression residuals were examined to confirm fulfilment of assumptions. The results indicated that skull and dental arcade variables yield little information about the size and physique of an individual but have some relevance for group comparisons. Long bone widths are moderately and equally correlated with height and weight. Lower limb bone widths are better correlated with body weight than those of the upper limb. The best predictive variables for body weight are the minimal ‘area’ of the lumbar vertebrae (L1–L4) and regressions involving various bone widths. It is confirmed that height correlates well with the lengths of long bones, but the prediction is improved by the addition of a width from the same bone as a second independent variable. Lower limb long bones are not better predictors of height than upper limb ones. The body mass index is best predicted from the available measurements by a regression with the minimal ‘area’ of L1 and the tibial length as independent variables. A lumbar vertebra, particularly L1, yields important information about height, weight and the body mass index. The patella width is, in part, a marker for mesomorphy. Reconstruction of the ankle to enable the width across the malleoli to be measured would yield useful estimates of weight and height. The findings have been applied to the Paviland femur. A provisional male attribution has been made. The derived height is similar to that of tall modern White men. The physique was probably mesomorphic and ectopenic, the physique of strength rather than mobility, but the confidence intervals are very wide. To enable this sort of assessment to be made more confidently, the study requires replication substituting the precision of magnetic resonance imaging for the relative imprecision of skeletal measurements taken directly from the intact body. Copyright © 1999 John Wiley & Sons, Ltd.  相似文献   

4.
Bergmann's and Allen's Rules predict a relationship between climate and morphology as a thermoregulatory adaptation. These ecogeographic principles predict/explain change in the ratio of body surface area to body mass in hot and cold climates. With regard to Allen's Rule we would expect short and relatively broad limb proportions. Such adaptation should be observable in skeletal architecture. This paper provides a test of Allen's Rule using osteometric data for the second metacarpal from the Sadlermiut of Southampton Island, Northwest Territories, Canada. Following adjustment for body size, ANOVA by sample and sex shows the Sadlermiut second metacarpal to be shorter, wider at the base and deeper at the distal metacarpophalangeal joint, in comparison to an historic sample of European settlers. This pattern of difference suggests an Inuit hand with a thermally adapted morphology, viz. a large mass relative to surface area. Copyright © 1999 John Wiley & Sons, Ltd.  相似文献   

5.
Trinkaus [Trinkaus, E., 2005. Anatomical evidence for the antiquity of human footwear use. J. Archaeol. Sci. 32, 1515–1526] provided a comparative biomechanical analysis of the proximal pedal phalanges of western Eurasian Middle Paleolithic and Middle Upper Paleolithic humans, in the context of those of variably shod recent humans. The anatomical evidence indicated that supportive footwear was rare in the Middle Paleolithic but became frequent by the Middle Upper Paleolithic. Based on that analysis, additional data are provided for the Middle Upper Paleolithic (∼27,500 cal BP) Sunghir 1 and the earlier (∼40,000 cal BP) Tianyuan 1 modern humans. Both specimens exhibit relatively gracile middle proximal phalanges in the context of otherwise robust lower limbs. The former specimen reinforces the association of footwear with pedal phalangeal gracility in the Middle Upper Paleolithic. Tianyuan 1 indicates a greater antiquity for the habitual use of footwear than previously inferred, predating the emergence of the Middle Upper Paleolithic.  相似文献   

6.
Cross-sectional growth data were obtained from the skeletal remains of non-adults from the Raunds Anglo-Saxon site. Standard measurements of the diaphyseal lengths of the long bones of the upper and lower limbs and the maximum breadth of the ilium were recorded in order to construct skeletal growth profiles (SGP). In addition regression equations were used to estimate diaphyseal length from proximal and distal shaft widths, and epiphyseal breadth data for fragmentary remains. The skeletal measurements were then plotted against age estimates determined by the dental formation standards of Moorrees, Fanning and Hunt, and Anderson, Thompson and Popovich. The growth data were compared with sixth to seventh century German, ninth century Slavic and modern Caucasian data. With the exception of the ancient Slavic material, the Anglo-Saxon remains demonstrated the smallest rates of growth. Diaphyseal ageing curves derived from the Anglo-Saxon sample were tested for applicability on the non-adult cohorts of the Berinsfield and Exeter Anglo-Saxon/Early Medieval samples. Differences were observed between diaphyseal age as determined from the skeletal growth profiles for Raunds and calcification age assessed for individuals within the test samples. It is proposed that variation in long bone growth as well as dental age confounds consistent and reliable ageing of skeletal remains based on diaphyseal length. Assessment of changes in health and evaluation of methodological problems inherent to studies of skeletal growth from archaeological populations are discussed. Population comparisons for changes in general health are recommended over individual assessments.  相似文献   

7.
Fifty-eight dental calculus samples from medieval and post-medieval skeletons from Vitoria, Spain, and a single sample from an Alaskan Inuit were tested for stable carbon and nitrogen isotope compositions. There was sufficient carbon and nitrogen concentrations to obtain δ13C and δ15N values, and the samples from Spain produced results that were replicable and comparable to European isotope values based on bone collagen collected from literature sources. The Alaskan Inuit calculus sample yielded a δ15N value of +17.5‰, well beyond the range of the Spanish samples, but consistent with literature data for modern Greenlandic Inuit consuming a diet rich in marine food. There are several potential sources for carbon and nitrogen in calculus. The results of this study yield stable isotope values consistent with those obtained from other biomaterials used as isotope proxies for paleodietary research, including bone collagen, hair, and fingernails, although further work is necessary to verify the fidelity of calculus as an isotope proxy. Many studies in bioarchaeology are precluded by curatorial concerns regarding the destructive analysis of primary biomaterials. However, calculus is an “add-on”, or secondary biomaterial, that is not an integral part of the dental or skeletal system. Hence, its consumption during analysis is technically not destructive. Therefore, isotope analysis of dental calculus may provide a potential new avenue for paleodietary analysis where the use of other primary biomaterials is precluded.  相似文献   

8.
Differences in adult male and female activity patterns may influence levels of sexual dimorphism in physical dimensions, including the cross‐sectional shape of long bone diaphyses. Previous studies of archaeological populations have demonstrated significant differences in diaphyseal shape between males and females. In this study, dimorphism in external diaphyseal shape of upper and lower limb bones (reflected in indices of external diaphyseal diameters), and bilateral asymmetry in these indices, were examined in two medieval populations: Muslim Écija (Spain) and Anglo‐Saxon Great Chesterford (UK). Attempts were made to relate observed patterns to documentary and other osteological evidence for differences in male and female activity patterns. While few significant differences in upper limb bone cross‐sectional shape were observed in either population, significant differences in shape were found in the lower limb diaphyses at Écija at the femoral midshaft and tibial foramen and midshaft levels, and at the tibial midshaft for Great Chesterford. Comparison with published data suggests that these differences are marked for Écija, and perhaps fairly high for Great Chesterford compared with other populations with an agriculture‐based economy. This is consistent with documentary and osteological evidence suggesting marked gender differences in behaviour in medieval Muslim Spain. No significant differences in bilateral asymmetry were found, but the effects of small sample size cannot be ruled out. Copyright © 2008 John Wiley & Sons, Ltd.  相似文献   

9.
Over the past four decades several workers have argued that the orientations of striations on the labial surfaces of the front teeth of Middle and Late Pleistocene fossils attributed to Neanderthals and antecedent archaic hominids can tell us something about handedness. These researchers have suggested that such scratches result from stone tools scraping across the incisors during ‘stuff‐and‐cut’ behaviours, and that they reflect directions of hand movements and handedness. In this study, we examine comparable wear features on the incisors of anatomically modern humans to determine whether striation angle may reflect handedness. The teeth from 66 individuals representing four groups with known differences in subsistence practices (Aleut, Arikara, Illinois Bluff, and Puye) were examined. Photomicrographs were made of the maxillary central incisor labial surfaces of all individuals at 56× magnification. These photomicrographs were scanned to image files, and orientations of all visible wear striations were measured using a semi‐automated image analysis procedure. Approximately three‐quarters of all labial surfaces showed a preferred striation orientation. Of those, most had preferred apico‐cervical or vertical striation orientations, independent of cultural affiliation. Few showed the tendency toward diagonally‐oriented scratches expected if handedness is related to scratch orientation. We found no evidence for ‘stuff‐and‐cut’ striations, despite the fact that at least one of the groups studied—the Aleut—are documented to have used this behaviour. In sum, there is evidently no relationship between striation orientation and handedness in the groups studied. It is, therefore, unlikely that labial scratch orientation in Middle and Late Pleistocene hominids reflects handedness if they used their front teeth in a manner comparable to that of any of the anatomically modern groups considered here. Copyright © 1999 John Wiley & Sons, Ltd.  相似文献   

10.
ABSTRACT

Today, one in four Inuit live outside of Inuit Nunangat, the Arctic territory incorporating the four northern Inuit land claims regions. With the majority of those Inuit living in cities, the issue of urban Inuit populations is becoming a more prominent focus for policy debate. In this article, I argue that urban Inuit mobility is a historically complex and multilayered phenomenon embedded in, not detached from, the major processes of social transformation affecting Inuit across the Canadian Arctic from the mid-twentieth century on. I use this analysis to explore the diverse formulation of Inuit-specific policy initiatives in Montreal, a task that, I show, is made easier by the Montreal Inuit radio show called Nipivut.  相似文献   

11.
Long distance trade has been attributed important social and economic roles in the pre-colonial south-central Andes, but how these trade networks were operated and organised, and the roles played by different populations and social groups (e.g. elites), remain uncertain. This study aims to offer new perspective on these questions through biomechanical analyses of human skeletal remains from a probable key site in these networks, San Pedro de Atacama (SPdA). Groups that were more intensively involved in long distance trade are expected to have been more habitually mobile, and thus to show greater robusticity and less circular lower limb bone cross-sections. Lower limb biomechanical properties of elite and non-elite Middle Horizon groups (MH, AD 500–1000) were compared with subsequent transitional MH-Late Intermediate Period (LIP, AD 1000–1450) and LIP groups from SPdA, and with LIP groups from Pica-8 and the Azapa Valley. The results indicate that MH populations from SPdA had less robust lower limbs and were by inference less mobile than their successors, with no differences between elite and non-elite, while robusticity was elevated in the MH-LIP transition group. Alternative explanations for the results, such as changes in herding activities, cannot be entirely discounted based on current evidence, but the results are consistent with hypotheses that SPdA may have served as a hub on long distance trade networks during the MH, before residents became more actively involved in long distance trade following the collapse of key links with the Tiwanaku polity. The results also indicate similar levels of robusticity among LIP populations at SPdA, Pica-8 and in the Azapa Valley, implying they may have been involved in trading activities to a similar extent, and perhaps to a greater extent than SPdA MH groups, as regional intergroup relationships changed.  相似文献   

12.
Palatine (PT) and mandibular torus (MT) have long been of interest to dental researchers and anthropologists, but their aetiology remains unresolved. Some combination of genetic and environmental factors influences their expression, but the relative role of each remains contentious. Previous research has shown that the Greenlandic Norse exhibit exceptionally high frequencies and pronounced expressions of PT and MT. In this regard, they are significantly different from genetically related medieval Scandinavian populations, so environmental factors have to be considered. An earlier study that estimated stable carbon and nitrogen isotope compositions for a Greenlandic Norse sample makes it possible to compare directly PT and MT expression with the relative degree of marine protein intake. For comparative purposes, parallel observations were made on a Greenlandic Inuit sample. Some researchers suggest the intake of marine resources could impact bone development, including torus expression, but our analysis found no significant correlation between PT or MT expression and δ13C and δ15N values in the Norse. In the Inuit, PT expression also showed no relationship to stable isotope compositions. MT size in the Inuit did, however, show a significant inverse relationship with δ13C and δ15N values. As MT size goes up, stable isotope compositions go down. Compared with contemporary European populations, the Greenlandic Norse show very positive isotope compositions, but the Inuit, with their high protein‐high fat diet, show significantly higher δ13C and δ15N values than the Norse. It is unclear how this relates to the contrasting findings for MT size and stable isotope values. Copyright © 2016 John Wiley & Sons, Ltd.  相似文献   

13.
Temporal changes in average stature are often used as a measure of a past population's adaptation, or lack of it. Traditionally, stature estimates have been calculated using formulae derived from limb proportions of cadavers. However, many authors have noted the problem of regional or population variation in body proportions of such reconstructed ratios. Before differences in stature can be attributed to environmental adaptation, ‘ethnic’ or population differences in limb ratios must be taken into account. The present paper calculates the stature of a medieval Norwegian skeletal sample using archaeological plan femur length and dry bone femur length. The author presents a variety of formulae and compares the stature derived from these calculations to the stature derived from archaeological plans. The Trondheim statures are then compared to stature reconstructions of other contemporary populations.  相似文献   

14.
The domestic dog fulfills many functions for their human companions, such as hauling, guarding, and protection. Consequently, humans have taken dogs to nearly every corner of the globe. Recent translocations of Western dogs stemming from the Victorian era dog fancy have erased some of the ancient genetic signatures of these earlier migrations. Here we used DNA of ancient and modern dogs from the North American Arctic of Alaska and Greenland to assess their genetic continuity in time and space. We successfully sequenced 23 archaeological (ca. AD 1250–1910) and recent (ca. AD 1930–1990) surface-collected Canis bone and tooth samples and compared them to 51 modern Inuit Sled Dogs and to published sequences of modern Alaskan Malamutes (and additional Inuit Sled Dogs) to test for evidence of lineage replacement or genetic continuity through time. Ancient samples from Alaska and Greenland and modern sequences from Greenland all contained a high frequency of haplotype A31, which was previously described only in modern North American Arctic dogs. Thus, A31 was a common thread tying the entire North American Arctic together prior to European colonization and, in the Eastern Arctic, indicates genetic continuity between past and present dogs as well. However, A31 is rare in modern Alaskan dogs, consistent with post-colonization replacement by Eurasian matrilines.  相似文献   

15.
This paper explores the geographic and environmental context of the Southern Dispersal Route, which has been proposed as a migratory route for Homo sapiens from East Africa to Australasia during oxygen isotope stage (OIS) 4 (71–59 kyr). A series of assumptions and constraints garnered from modern hunter-gatherer observations are used to build a model of coastal foragers, which is then integrated with high-resolution physiographic analyses to produce a potential dispersal route along the coastline of the Indian Ocean. Paleoenvironmental conditions that may have supplied critical resources or served as obstacles to human colonization are identified and discussed in regards to human subsistence, the speed of migration, and demographic expansion. These factors suggest that rapid dispersals along coastlines and river valleys would have occurred upon the initial expansion out of Africa, but slowed as populations expanded demographically into South Asia and the Sunda Shelf. This also suggests that archaeological signatures relating to the earliest modern Homo sapiens are more likely to be recovered in South Asia.  相似文献   

16.
The present bioarchaeological study examines the external diaphyseal geometric properties of humeri, radii, femora and tibiae of the Classic period skeletal population of Xcambó, Yucatan, Mexico. The diaphysial proportions are evaluated using a biomechanical approach together with data from the material context and other osteological information. Our intent is to provide new answers to questions concerning lifestyle, domestic labour division and subsistence strategies of this coastal Maya settlement that was inhabited from the Late and Terminal Preclassic (300 BC–350 AD) to the Postclassic Period (900–1500 AD). Our results provide evidence for a marked sexual division of labour when compared with values from contemporaneous inland populations. The overall male and female loading patterns differ remarkably in terms of form and in bilateral comparison. A high directional asymmetry in the upper limbs is evident among males, a condition related to maritime transportation and trading activities. On the other hand, female upper limbs are characterized by very low side differences. Forces on the arms of women were probably dominated by food processing, in particular the grinding of grains or seeds. In the lower limbs, males show significantly higher anteroposterior bending strengths, which can be explained by greater engagement in transportation tasks and carrying heavy loads. In the course of the Classic period (350–900 AD), diachronic changes affect the male sample only, which suggests a shift of occupational pattern and physical demands. This shift, in turn, reflects Xcambó's changing role as the centre of a densifying settlement area and its place in the trading activities of northern Yucatan. Other topics of discussion relate to general regional trends and local prehispanic subsistence strategies. Our conclusions emphasize the value of geometric long bone analysis in the reconstruction of activity patterns and lifestyles in ancient coastal settlements. Copyright © 2007 John Wiley & Sons, Ltd.  相似文献   

17.
Osteoarthritis is among the most common pathological conditions in skeletal collections and is the most frequent musculoskeletal disorder in contemporary populations. Jurmain ( 1991 ) has previously published in this journal a brief review of skeletal perspectives on osteoarthritis. Subsequent studies by osteologists and medical researchers have added considerably to understanding of the aetiology and patterning of osteoarthritis. Thus, it is timely to present an updated review that expands and supports conclusions discussed in the earlier review. In short, osteoarthritis aetiology is multifactorial, with age being the main influence on the onset and severity of osteoarthritis. Genetic influences also play a large role in the severity of osteoarthritis, especially in the lower limbs. Weight, although playing a significant role for modern populations, seems to have had very minimal effects on prehistoric populations. Sex differences may often be a consequence of hormones, body size and anatomy, rather than activity related. Finally, intense activity starting at a young age still may influence osteoarthritis, especially in the upper limbs. Future directions discussed include within‐body comparisons, animal studies, and examining patterns in large populations. Copyright © 2007 John Wiley & Sons, Ltd.  相似文献   

18.
Several lines of archaeological and human palaeontological data from the Moravian earlier Gravettian (or Pavlovian) indicate a high level of mobility among these earlier Upper Palaeolithic human populations. The congenitally abnormal young adult Dolnı́ Věstonice 15 human remains support this pattern. Dolnı́ Věstonice 15 suffered serious developmental abnormalities, most likely as a result of the X-linked dominant form of chondrodysplasia calcificans punctata, that included pronounced infantile and minor later childhood developmental defects of dental enamel (enamel hypoplasia), asymmetrical curvature and angular deformities of the proximal femora, asymmetrical shortening of the femora, a diaphyseal deformity of the distal right humerus, a probable healed left ulnar fracture with associated deformity of the radial diaphysis, elevated length asymmetries of the forearms, and an unusual pattern of anomalies of tooth position and number. These are combined with trivial intervertebral articular degenerations, plus localized pronounced osteoarthritis of the right glenohumeral joint and right second and fifth ray hand proximal interphalangeal joints. Yet, there is little asymmetry of the lower limbs distal of the mid-femora, and both diaphyseal hypertrophy of the humeri, femora and tibiae and relative arm and leg muscle attachment size and moment arms indicate a level of skeletal hypertrophy average to high for an earlier Upper Palaeolithic human. The severe upper limb osteoarthritis may indicate a pattern of pronounced and repetitive carrying or dragging of heavy loads. These data indicate the ability of the population to keep a severely affected individual alive through development, yet they also emphasize the necessity for all individuals to participate actively in the elevated mobility and overall activity levels of these earlier Upper Palaeolithic human populations.  相似文献   

19.
The anatomy of a new articulated enantiornithine bird skeleton from the Late Cretaceous of Patagonia clearly indicates a capacity for powered flight, approaching that of modern birds. Enantiornithines possess some of the synapomorphies of the Ornithurae, although they retain plesiomorphic states for many other characters, mainly in the hind limb. Such a mosaic character combination suggests a sister-group relationship between Enantiornithes and Ornithurae. Derived features of the pectoral girdle are here considered as diagnostic for a major avian clade, the Ornithopectae, comprising all known birds other than Archaeopteryx. The combination of derived and primitive traits in the fore and hind limbs and their girdles in early ornithopectines reflects mosaic evolution, with flight-related modifications of the fore limb and pectoral girdle preceding those in the hind limb and pelvic girdle.  相似文献   

20.
Interest in sensations from removed body parts other than limbs has increased with modern surgical techniques. This applies particularly to operations (e.g., gender-changing surgeries) that have resulted in phantom genitalia. The impression given in modern accounts, especially those dealing with phantoms associated with penis amputation, is that this is a recently discovered phenomenon. Yet the historical record reveals several cases of phantom penises dating from the late-eighteenth century and the early-nineteenth century. These cases, recorded by some of the leading medical and surgical figures of the era, are of considerable historical and theoretical significance. This is partly because these phantoms were associated with pleasurable sensations, in contrast to the loss of a limb, which for centuries had been associated with painful phantoms. We here present several early reports on phantom penile sensations, with the intent of showing what had been described and why more than 200 years ago.  相似文献   

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