An experimental study of large mammal bone modification by crocodiles and its bearing on the interpretation of crocodile predation at FLK Zinj and FLK NN3

The taphonomic signature of crocodiles as agents of bone modification has been previously identified by specific tooth mark types (e.g., bisected pits) and by a conspicuous presence of these marks: more than 75% of bones modified by crocodiles bear at least one of these distinctive marks. Therefore, crocodile tooth-marking would be notably prevalent in bone assemblages resulting from crocodile predation and active scavenging. The present study contributes to refine this diagnosis by showing greater variability of these types of marks, a different degree of tooth-marking intensity, and a somewhat different distribution of tooth-marked elements according to skeletal parts from previous experiments with crocodiles. Some of these differences are due to different experimental variables and conditions and this highlights the need to understand behavioral variability in crocodile ecological settings. This variability in crocodile tooth-marking probably results from several as-yet-inadequately measured behavioral and ecological factors, such as intensity of feeding competition and differences between male and female crocodile feeding behaviors, among others. Furthermore, this study also contributes to a better definition of the microscopic criteria that can be used to distinguish crocodile-inflicted marks from other types of bone surface modifications. In light of these and previous experimental frameworks, we reevaluate the application of these analogs to modifications documented in hominin fossils from Olduvai Gorge (OH8 and OH35) and the resulting inferences about the hazard posed by crocodiles on the paleolandscape where FLK North North and FLK Zinj (Bed I) were formed. The taphonomic analysis also shows that Olduvai OH8 and OH35 were probably not preyed upon by crocodiles. It is concluded that no tangible evidence can be used to support the interpretation that OH35 was modified by crocodiles and that the overall presence of crocodiles in FLK North North and FLK Zinj was rather marginal, based on the virtual absence of crocodile-modified bones in both archaeofaunal assemblages.     

Ascertaining an agent: using tooth pit data to determine the carnivore/s responsible for predation in cases of suspected big cat kills in an upland area of Britain

The identification of the involvement of a particular carnivore in the modification of bone assemblages concerns a number of fields of research including archaeological and palaeontological enquiry. Taphonomy provides a methodology by which bone assemblages can be analysed and interpreted and this is more often undertaken with archaeological or palaeontological assemblages. A taphonomic analysis is undertaken here in order to determine the perpetrator of predation attacks on domestic stock from a modern-day setting. Recently reported techniques using tooth marks preserved on bone surfaces made by known carnivores are successful at determining some class sizes of predators and are used here to determine the perpetrator(s). Although a class size of carnivore is readily identified by this methodology, a particular carnivore taxon is not. Tooth morphology and dental configuration are reported here as better criteria for identifying a particular taphonomic agent. Tooth pit dimensions are used here to identify the class size of carnivores involved, and tooth morphology and cusp spacing to suggest a medium sized felid and fox as taphonomic agents. The identification of the medium-sized felid may support observations and reports of alleged “big” cat kills in the area. The study has important implications for the interpretation of fossil sites where felids may have been involved in the modification of animal carcasses but are archaeologically invisible in terms of their fossil remains.     

Variation in tooth mark frequencies on long bones from the assemblages of all three extant bone-collecting hyaenids

Tooth mark frequencies on long bones are examined from the assemblages of all three extant bone-collecting hyaenids. Comparisons are made with a recent study examining tooth mark frequencies and possible sources of variation from a single spotted hyaena (Crocuta crocuta) assemblage (Faith, J.T., 2007. Sources of variation in carnivore tooth-mark frequencies in a modern spotted hyena (Crocuta crocuta) den assemblage, Amboseli Park, Kenya. Journal of Archaeological Science 34 (10), 1601–1609). The factors that may influence tooth mark frequencies are fragment size, fragments from different sized animals, region of skeletal element and bone density. All four factors are examined in the present study and compared across species and with previous results. The results indicate that there is a great deal of variation in tooth mark frequencies not only between the species but also from the same species.     

A cautionary note on the use of captive carnivores to model wild predator behavior: a comparison of bone modification patterns on long bones by captive and wild lions

A study with wild lions in Tarangire National Park (Tanzania) and with captive lions in Cabárceno Reserve (Spain) has yielded two different bone modification patterns, probably as a result of the differences in environmental contexts. Captive lions have modified bones more intensively, both in the form of total number of tooth-marked bones and number of tooth marks per tooth-marked bone, probably because of stereotypic behaviors. This emphasizes the importance of environmental contexts to understand carnivore behavior and their resulting bone modification patterns. It also shows that analogical models based on experiments carried out with captive carnivores may be biased and inadequate as proxies for wild carnivore bone modification behaviors.     

Absolute age and tooth eruption and wear sequences in sheep and goat: determining age-at-death in zooarchaeology using a modern control sample

This paper seeks to improve our understanding of the relationship between mandibular tooth eruption and wear sequences and absolute age in modern sheep and goat. While the relationship between tooth eruption and wear and absolute age has been investigated for primitive breeds in parts of the Old World, our pilot study extends the applicability of this research to modern breeds and to another geographic region (North America). A modern control sample of two species of domestic caprines – Ovis aries (sheep) and Capra hircus (goats) from Manitoba, Canada – are used to evaluate the two major systems for analyzing mandibular tooth wear and eruption (Payne and Grant) and for assigning an absolute age-at-death. In order to compare the two systems, an absolute age for Grant's Mandibular Wear Stages was established for the first time, previously demonstrated only for Payne's system. The analysis confirmed that there is a very high correlation between the known age and estimated age based on tooth eruption and wear. However, Payne's system generated far coarser age-at-death profiles, which lowers its utility in constructing harvest profiles. Grant's system is recommended for ageing be utilized since it can yield more sensitive age estimations. In conclusion, it is possible to utilize the modern caprine breeds to establish the relationship between tooth eruption and wear and absolute age to reconstruct age and season of death of archaeological samples.     

Aging mountain gazelle (Gazella gazella): refining methods of tooth eruption and wear and bone fusion

This paper refines current methods for aging mountain gazelle (Gazella gazella), one of the most common species hunted by prehistoric humans in the southern Levant. Most importantly, we present a new tooth wear scheme for aging the lower deciduous fourth premolar (dP₄), the lower permanent molars (M₁, M₂, M₃) and the lower fourth premolar (P₄) of the gazelle. Broader wear stages for complete mandibles and individual teeth and bone fusion stages are also provided. Finally, a comparison of the tooth wear and bone fusion data indicates that the two methods can be linked at the important transition when a gazelle reaches adulthood (ca. 18 months of age). The completion of the fusion of the gazelle skeleton corresponds directly to the replacement of the dP₄ by the P₄ and the beginning of wear of the M₃. The coincidence of these aging indicators allows for direct comparison of gazelle bone fusion and tooth wear data.     

Towards a distinction between digested and undigested glume bases in the archaeobotanical record from Neolithic northern Greece: A preliminary experimental investigation

Abstract

This study investigates the possibility of distinguishing digested from undigested glume wheat chaff in the archaeobotanical record. To this end the contents of dung pellets collected from a goat fed on einkorn spikelet forks and glume bases are analysed and the derived einkorn chaff examined macroscopically with the aid of scanning electron microscopy. The modern dung pellets contained recognisable glume bases which, compared with undigested ones, demonstrated a ‘rugged’ surface. When these dung pellets were charred, glume bases were still preserved in them. The experimental pellets show that dung can contribute glume bases in archaeobotanical assemblages and that these glume bases may be distinguishable from those derived from glume wheat dehusking by-products used as fuel. These results need to be further explored on archaeobotanical materials. Moreover, further experimentation with different charring regimes, as well as the examination of more experimental specimens is needed.     

Change and variability in Plio-Pleistocene climates: modelling the hominin response

Research into the links between climatic change and hominin evolution has generated numerous hypotheses. In recent years, methodological refinement of, and increased research effort directed towards, reliable proxies for palaeoclimatic change have provided a growing body of data with which to test such hypotheses. Whilst many archaeologists are aware of these data, few are cognizant of the wealth of techniques developed by theoretical biologists over the last half-century to explicitly address the evolutionary consequences of adaptation to temporally heterogeneous environments. The current paper expands and adapts one such technique for use with empirical data, and applies it to a global palaeoclimatic record spanning the last five million years, in order to discern the potential impact of environmental heterogeneity on hominin evolution during this period. Of particular interest are the contributions of climatic change, associated with directional selection, and climatic variability, associated with selection for phenotypic plasticity. At this macro-scale, results suggest an early peak in selection for plasticity at approximately 2–2.7 mya, combined with three major shifts in directional selection at approximately 3.3–3.4, 1.4–1.5, and 0.5–0.6 mya. These results are employed to relate the fossil and archaeological records to a number of environmental hypotheses of human evolution. In particular, it is argued that the origins of the genus Homo and the spread of Oldowan technology are associated not with a major turnover pulse, but with a period of selection for phenotypic plasticity.     

Articulating minority nationhood: cultural and political dimensions in Québec's reasonable accommodation debate

Given their precarious position within larger states, national minorities cannot rely on federal governments to affirm their nationhood. Moreover, insofar as nationhood is predicated on a shared history, language and culture, immigrants place additional strains on the maintenance of national distinctiveness and the political claims that derive from it. In 2006–2007, following a series of confrontations over religious practices in the public sphere, Québec's provincial government appointed the Bouchard–Taylor Commission to investigate avenues for the accommodation of immigrant‐related cultural and religious differences. While it failed to generate policy, the commission did provide a discursive space for the (re)assertion of Québécois nationhood. Analysing the production of national identity in newspaper debates of the Bouchard–Taylor report, we offer an alternative to the ethnic–civic paradigm in nationalism theory. Rather than treat ethnic and civic as two separate ends of a single continuum, we conceptualise a relationship between two dimensions: one of culture and one of politics. We show that in contemporary articulations of Québec national identity, the prerequisites of political membership derive their meaning from a productive tension between blood‐based and adoptive conceptions of national culture.     

Archaeobotany,vine growing and wine producing in Roman Southern France: the site of Gasquinoy (Béziers,Hérault)

Unequivocal evidence of Roman vine cultivation and wine making is provided from studies of combined archaeological remains from the site of Gasquinoy (Southern France). Waterlogged and charred plant material (fruits/seeds/wood) collected from wells located in the close vicinity of cultivated fields and wine making establishments confirms the local significance of this activity. The results offer insights on particular aspects of wine production (‘traumatic’ treading of grapes and straining) and provide evidence of secondary agricultural activities such as cereal production and fruit tree cultivation. The potential use of monocotyledonous stems such as Arundo/Phragmites in the farming system is discussed.     

Solubility of corundum + kyanite in H2O at 700°C and 10 kbar: evidence for Al‐Si complexing at high pressure and temperature

The solubility of the assemblage corundum + kyanite in H₂O was determined at 700°C and 10 kbar, using a piston‐cylinder apparatus and rapid‐quench/fluid‐extraction techniques. Weighted mean concentrations of total Al and Si were 5.80 ± 0.03 mmol kg^?1 H₂O and 0.308 ± 0.003 mol kg^?1 H₂O, respectively (1σ errors). The Al concentration is nearly five times higher than that of corundum solubility in pure H₂O. This difference is interpreted to arise from complexing between Si and Al to form HAlSiO_4,aq species. Charged or more polymerized species are also possible, but their abundance cannot be constrained based on these experiments. Assumption of a single aqueous aluminosilicate complex permits calculation of the thermodynamic consequences of Al‐Si interaction in high‐pressure fluids, as well as phase diagrams for the system Al‐Si‐O‐H. Formation of Al‐Si complexes leads to a large increase in dissolved Al with increasing Si in solution, such that Al concentration in equilibrium with kyanite + quartz is predicted to be 7.1 mmolal, higher than with kyanite + corundum. The elevated concentration of Si in deep‐crustal and mantle aqueous fluids suggests that Al must readily be dissolved and transported by Al‐Si complexing in high‐pressure metamorphic and metasomatic environments. The results provide a simple explanation for the common observation of kyanite + quartz segregations in eclogites and Barrovian metamorphic rocks.     

Paleoindian subsistence strategies and late Pleistocene paleoenvironments in the northeastern and southwestern United States: a tooth wear analysis

Paleoindians of North America entered a continent undergoing rapid climatic and environmental changes. This paper is a preliminary contribution toward obtaining a better picture regarding how climate and environmental change might have impacted the first settlers of North America. The Paleoindian sites we analyzed are, from the oldest to the youngest, Ingleside (Texas), Blackwater Draw Locality No. 1 (New Mexico), Hiscock Site (New York), and Plainview Quarry (Texas). Paleoenvironmental reconstruction involves identifying the dietary traits of ungulate species that might reflect the environmental conditions where they were living, and also where they might have been hunted by Paleoindians. Such an approach is realized through tooth microwear and mesowear analyses. Results indicate that a variety of food resources were available for the ungulates at these sites as well as the likely presence of mosaic environments around these sites which allowed Paleoindians to exploit resources in a large variety of habitats. The application of a method that allows for the estimation of the duration of occupation at archaeological sites reveals that results for Blackwater Draw and Hiscock Site indicate a long-term occupation of probably several months. However, during short events, Paleoindians were most likely hunting herds of horse and bison when these prey were available near the site. Results indicate that Plainview Quarry was likely used only for short-term occupations, with large game hunting focusing on bison. These patterns identified at the archeological sites studied are related to the fact that Paleoindians follow a high-technology forager model and frequently shifted their territory depending on the composition and distribution of the large mammal fauna.     

Sr/Sr variability in Puerto Rico: geological complexity and the study of paleomobility

The temptation to use biogeochemical techniques to resolve issues of paleomigration is evident and well intentioned. Knowledge of radiogenic strontium isotope baselines in a region of interest is a sine qua non of such archaeological studies of paleomobility. Here, we present the first detailed study of baseline ⁸⁷Sr/⁸⁶Sr values for the island of Puerto Rico. The high degree of ⁸⁷Sr/⁸⁶Sr variability present in this corpus of modern Puerto Rican bedrock and terrestrial malacological samples (0.70406–0.70909) is a testament to the complex geology of that island. This diversity of ⁸⁷Sr/⁸⁶Sr values makes parsing issues of origin a difficult and highly contingent task. Given these complexities, regional studies seeking to assess paleomigration by such isotopic means should proceed with a great deal of caution.     

Residential mobility and social identity in the periphery: strontium isotope analysis of archaeological tooth enamel from southeastern Arabia

Strontium isotope analysis represents an effective means of assessing mobility and reconstructing geographic residence patterns in archaeological populations. This biogeochemical technique was utilized to test the hypothesis that burgeoning interregional exchange networks and the occurrence of exotic grave goods in local tombs would correspond with a highly mobile population and a considerable immigrant presence during the Umm an-Nar (2700–2000 BC) period in the UAE. This region has been considered peripheral relative to larger civilizations in Mesopotamia and the Indus Valley, but played an important role as a major supplier of copper for the Persian Gulf. Individuals (n = 100) from six monumental Umm an-Nar tombs (Mowaihat; Tell Abraq; Umm an-Nar Island I, II, V; Unar 1) were selected to evaluate the geographic origins of tomb members. Mean ⁸⁷Sr/⁸⁶Sr ratios from local individuals interred at Mowaihat (0.708863 ± 0.000014; 1σ, n = 12), Tell Abraq (0.708873 ± 0.000020; 1σ, n = 27), Umm an-Nar Island (0.708902 ± 0.000079; 1σ, n = 33), and Unar 1 (0.708805 ± 0.000065; 1σ, n = 25) all display little isotopic variability, indicative of a population that was not highly mobile. However, coupled with archaeological evidence, three immigrants from Tell Abraq (n = 2) and Mowaihat (n = 1) identified by deviant strontium values suggest that this region was actively engaged in interregional interaction. Despite claims that these tombs acted as visible markers of territoriality legitimized by local ancestors buried within them, the presence of non-locals suggests that as commerce became increasingly important, definitions of kinship and social identity may have become more flexible to better meet the needs of the local community and those with whom they interacted.     

The Natufian: Settlement variability and economic adaptations in the Levant at the end of the Pleistocene

The relationship between current interpretations of Natufian settlement and subsistence and available archaeological data are examined in light of recent research, particularly in Jordan. Regional variability in adaptive strategies is discernible, particularly between forest and coastal sites versus steppe and desert sites. Greater evidence of plant processing and more intensive occupation characterize settlement in the former, although year-round occupation has yet to be conclusively demonstrated. Patterned variability also exists between two classes of steppe and desert area settlements. One set of steppe and desert sites is characterized by a broad range of activities and moderate settlement permanence and activity intensity, while less permanent occupation and more specialized activities focused primarily on hunting typify the other set of sites. Evidence for food production in the Natufian is examined and, although the domestication process may have begun, no morphological evidence exists for the domestication of plants or herd animals. Finally, worthwhile areas for future research are outlined.     

Seasonality and season of birth in early Eneolithic sheep from Cheia (Romania): methodological advances and implications for animal economy

Cheia (early 5th millennium cal BC) is a prehistoric village in the Dobrodgea province in Romania. Its occupation is attributed to the Early Eneolithic period or Hamangia III phase. The exploitation of animal resources is heavily dominated by husbandry. Although cattle are dominant, they were complemented by caprines, mainly sheep, exploited for tender meat (as highlighted by a 6–12 months age class peak in the mortality profile). Sheep reproduction patterns were investigated through stable isotope analyses in order to characterize the annual rhythms of slaughtering for tender meat acquisition while informing a more general picture on sheep demographical management and animal husbandry at the settlement. Results from δ¹⁸O analysis on second and third molar enamel were modeled and compared with modern reference populations. Sheep births took place over less than four months, from late winter to early summer. From this it could be concluded that tender meat could be provided most of the year, excepted over short period in early summer. Finally, comparison of datasets obtained on the M2 and M3 suggests that the M2 presents a more accurate representation of birth seasonality due to lower inter-individual variability in the chronology of tooth growth.     

Solubility of quartz in crustal fluids: experiments and general equations for salt solutions and H2O–CO2 mixtures at 400–800°C and 0.1–0.9 GPa

The solubility of quartz has been measured in a wide range of salt solutions at 800°C and 0.5 GPa, and in NaCl, CaCl₂ and CsCl solutions and H₂O–CO₂ fluids at six additional P–T conditions ranging from 400°C at 0.1 GPa to 800°C at 0.9 GPa. The experiments cover a wide range of compositions along each binary. At P–T conditions where the density of pure water is low (0.43 g cm^?3), addition of most salts produces an enhancement of quartz solubility at low to moderate salt concentrations (salt‐in effect), although quartz solubility falls with further decrease in X_H2O. At higher fluid densities (0.7 g cm^?3 and greater), the salt‐in effect is generally absent, although this depends on both the cation present and the actual P–T conditions. The salt‐in effect is most readily produced by chloride salts of large monovalent cations, while CaCl₂ only produced a salt‐in effect at the most extreme conditions of high‐T and low‐P investigated (800°C at 0.2 GPa). Under most crustal conditions, the addition of common salts to aqueous fluids results in a lowering of quartz solubility relative to that in pure water (salt‐out effect). Comparing quartz solubility in different fluids by calculating X_H2O on the basis that all salts are fully associated under all conditions yields higher quartz solubility in solutions of monovalent salts than in solutions of divalent salts, absolute values are also influenced by cation radius. Quartz solubility measurements have been fitted to a Setchenow‐type equation, modified to take account of the separate effects of both the lowering of X_H2O and the specific effects of different salts, which are treated as arising through distinct patterns of non‐ideal behaviour, rather than the explicit formation of additional silica complexes with salt components. Quartz solubility in H₂O–CO₂ fluids can be treated as ideal, if the solvation number of aqueous silica is taken as 3.5. For this system the solubility (molality) of quartz in the binary fluid, S is related to its solubility in pure water at the same P–T conditions, S^o, by: Quartz solubility in binary salt systems (H₂O–RCl_n) can be fitted to the relationship: where salt concentration mRCl_n is expressed as molality and the exponent b has a value of 1 except under conditions where salting‐in is observed at low salt concentrations, in which case it is <1. Under most crustal conditions, the solubility of quartz in NaCl solutions is given to a good approximation by: We propose that quartz solubility in multicomponent fluids can be estimated from an extended expression, calculating X_H2O based on the total fluid composition (including dissolved gasses), and adding terms for each major salt present. Our experimental results on H₂O–NaCl–CO₂ fluids are satisfactorily predicted on this basis. An important implication of the results presented here is that there are circumstances where the migration of a fluid from one quartz‐bearing host into another, if it is accompanied by re‐equilibration through cation exchange, may lead to dissolution or precipitation of quartz even at constant P and T, with concomitant modification of the permeability structure of the deep crust.     

A method for estimating edge length from flake dimensions: use and implications for technological change in the southern African MSA

This paper presents a method for estimating edge length from flake dimensions. Comparison of estimated values with data from measured flake edges demonstrates the robusticity of the method. Building on the work of Braun (Braun, D.R., 2005. Examining flake production strategies: examples from the Middle Paleolithic of Southwest Asia. Lithic Technology 30, 107–125; Braun, D.R., Harris, J.W.K., 2003. Technological developments in the Oldowan of Koobi Fora: innovative techniques of artifact analysis and new interpretations of Oldowan behavior, in Mora, R., de la Torre, I. (Eds.), Oldowan: Rather More than Smashing Stones. Treballs d’ Arqueologia 9, pp. 132–144) this estimate is used to generate an edge length to mass value for complete flakes—taken to be a proxy for flaking efficiency. It is shown that this value is useful in tracking the onset, variability and decline of a well known technological phase—the Howiesons Poort of South Africa. The results suggest that the Howiesons Poort was a time in which flaking efficiency was emphasised. Furthermore, comparison of upper limits of edge to mass values for different raw materials may provide an insight into changes in raw material selection.     

Age estimation by tooth/pulp ratio in canines by peri-apical X-rays: reliability in age determination of Spanish and Italian medieval skeletal remains

Estimation of age at death is an unavoidable step in the process of human identification, both in forensic practice and in the anthropological and palaeopathological study of skeletal remains. In several cases, in which medical or demographic records are completely lacking, a reliable estimation of the age at death becomes very important. Skeletal remains from archaeological contexts suffer from several biasing factors such as post-mortem changes, taphonomy and various burial practices depending on age, sex and social status of the deceased persons.     

Applying new approaches to modeling diet and status: isotopic evidence for commoner resiliency and elite variability in the Classic Maya lowlands

Classic Maya states were characterized by a high degree of socioeconomic stratification. This paper investigates the degree to which status, as defined by grave goods and tomb construction, influenced dietary patterns of elites and commoners throughout the Classic Period (200–900/1000 AD) of the southern lowlands. We compile a database (N = 102) of previously-published stable isotope ratios (δ¹³C collagen, δ¹³C apatite, and δ¹⁵N collagen) from Maya bone mineral and collagen, and interrogate these data through two new isotopic modeling techniques: a simple carbon isotope model ( Kellner and Schoeninger, 2007; Froehle et al., 2010) and a multivariate isotope model ( Froehle et al., 2012). We find that Maya elite diet varied significantly through time in terms of maize consumption and trophic level, while commoner diet remained remarkably stable. These findings provide new information relevant to studies of ancient Maya class structure and to studies of subsistence strategies of the pre-Columbian Americas.