The Middle Paleolithic of the East Mediterranean Levant

This paper reviews recent developments in geochronology, archaeology, and behavioral interpretations of the Middle Paleolithic Period (ca. 47–250 Kyr) in the East Mediterranean Levant. Neandertals and early modern humans both occupied the Levant during this period. Both these hominids are associated with the Levantine Mousterian stone tool industry and similar sets of faunal remains. The Levant has long been seen as preserving evidence for the origin of modern humans out of Neandertal ancestors. Recent radiometric dates for Levantine Middle Paleolithic contexts challenge this hypothesis. Instead, they suggest the evolutionary relationships between these hominids were far more complicated. Proposed models for Neandertal and early modern human coevolutionary relationships are examined. Intense competition between Neandertals and early modern humans for a narrow human niche may be the context out of which the Upper Paleolithic behavioral revolution arose.     

Repeat silica extraction: a simple technique for the removal of PCR inhibitors from DNA extracts

Polymerase chain reaction (PCR) inhibitors are often co-extracted with ancient DNA (aDNA) and when present make the analysis of aDNA difficult, if not impossible. In this study we review previous research on PCR inhibitors and techniques that address their co-extraction with DNA from sub-optimal samples. Additionally, we introduce a simple extraction technique, “repeat silica extraction,” that effectively removed PCR inhibitors from extracts of 7000–8000-year-old human skeletal remains from the Windover archaeological site in Florida and 700–2000-year-old human coprolites excavated from Fish Slough Cave in southern California. A series of tests on these same samples demonstrates that N-phenacylthiazolium bromide is largely ineffective, despite previously reported success using this compound as part of the DNA extraction process. We also describe a method for demonstrating the presence as well as successful removal of PCR inhibitors by use of a “positive aDNA control,” a test necessary to conclude that negative PCR amplification is the result of the absence of preserved DNA.     

The Emergence of Ornaments and Art: An Archaeological Perspective on the Origins of “Behavioral Modernity”

The earliest known personal ornaments come from the Middle Stone Age of southern Africa, c. 75,000 years ago, and are associated with anatomically modern humans. In Europe, such items are not recorded until after 45,000 radiocarbon years ago, in Neandertal-associated contexts that significantly predate the earliest evidence, archaeological or paleontological, for the immigration of modern humans; thus, they represent either independent invention or acquisition of the concept by long-distance diffusion, implying in both cases comparable levels of cognitive capability and performance. The emergence of figurative art postdates c. 32,000 radiocarbon years ago, several millennia after the time of Neandertal/modern human contact. These temporal patterns suggest that the emergence of “behavioral modernity” was triggered by demographic and social processes and is not a species-specific phenomenon; a corollary of these conclusions is that the corresponding genetic and cognitive basis must have been present in the genus Homo before the evolutionary split between the Neandertal and modern human lineages.     

Archaeological Evidence for the Emergence of Language,Symbolism, and Music–An Alternative Multidisciplinary Perspective

In recent years, there has been a tendency to correlate the origin of modern culture and language with that of anatomically modern humans. Here we discuss this correlation in the light of results provided by our first hand analysis of ancient and recently discovered relevant archaeological and paleontological material from Africa and Europe. We focus in particular on the evolutionary significance of lithic and bone technology, the emergence of symbolism, Neandertal behavioral patterns, the identification of early mortuary practices, the anatomical evidence for the acquisition of language, the development of conscious symbolic storage, the emergence of musical traditions, and the archaeological evidence for the diversification of languages during the Upper Paleolithic. This critical reappraisal contradicts the hypothesis of a symbolic revolution coinciding with the arrival of anatomically modern humans in Europe some 40,000 years ago, but also highlights inconsistencies in the anatomically–culturally modern equation and the potential contribution of anatomically pre-modern human populations to the emergence of these abilities. No firm evidence of conscious symbolic storage and musical traditions are found before the Upper Paleolithic. However, the oldest known European objects that testify to these practices already show a high degree of complexity and geographic variability suggestive of possible earlier, and still unrecorded, phases of development.     

Wild or domesticated: DNA analysis of ancient water buffalo remains from north China

Recent zooarchaeological studies on water buffalo (Bubalus sp.) remains from China and south Asia question the traditional view that water buffalo were first domesticated in Neolithic China over 7000 years ago. The results from several recent population genetic studies of modern domesticated buffalo (Bubalus bubalis) are not consistent with each other, placing the original center of buffalo's domestication in south Asia, southeast Asia, or China. This paper reports a study using an ancient DNA approach to analyze water buffalo remains from Neolithic sites in north China to investigate their affinities with modern domesticated water buffalo, and to shed light on the origin of modern domesticated water buffalo in China.A 169 bp fragment of D-loop mitochondrial DNA was successfully amplified and verified for 13 of 24 bone samples obtained from seven archaeological sites along the Wei River valley in Shaanxi Province, China. The bone samples which yielded positive DNA can be dated to 8000–3600 cal. BP. The phylogenetic analysis of the obtained DNA sequences along with modern water buffalo sequences indicated that the ancient water buffalos were not the direct ancestor of modern domesticated water buffalo. However, the phylogenetic analysis, along with BLAST searches of these ancient DNA sequences, did demonstrate their relatedness to water buffalo more so than to any other bovid species, confirming the existence of indigenous wild (but now extinct) water buffalo species (B. mephistopheles) in ancient China.The DNA analysis of these ancient remains failed to establish direct links between modern domesticated water buffalo (B. bubalis) and indigenous water buffalo (B. mephistopheles) from ancient China. If further DNA studies of more ancient remains from other regions of China confirm the observation of solely indigenous water buffalo species in ancient China, it would suggest modern water buffalo might not have been first domesticated in China.     

Possible evidence of bone tool shaping by Swartkrans early hominids

Ever since Dart (J. Phys. Anthrop. 7 (1949) 1) interpreted certain bones from Makapansgat as tools, scientific consensus has fluctuated as to whether some bone objects from early hominid sites should be interpreted as artefacts, or the result of non-human taphonomic processes, which are known to produce pseudo-bone tools morphologically similar to human modified or used artefacts. Here we present possible evidence of bone tool shaping from Swartkrans (Members 1–3; ca. 1.8–1.0 Mya). Four horncores and the proximal end of an ulna used as tools in digging activities also have facets covered by parallel spindle-shaped striations characteristic of grinding. Identification of these traces as possibly resulting from deliberate shaping or re-sharpening of the bone tools is based on the characterisation of the use-wear pattern and other taphonomic modifications observed on the Swartkrans bone tools. This interpretation is also supported by the study of the remainder of the horncores from Swartkrans, horncores from other southern African Plio-Pleistocene sites (Sterkfontein, Makapansgat, Gondolin), modern horncores affected by pre- and post-mortem modification, ethnographic, LSA, African Iron Age and experimental bone tools shaped by grinding. These data suggest that early hominids had the cognitive ability to modify the functional area of bone implements to achieve optimal efficiency.     

The taphonomy of fallow deer (Dama dama) skeletons from Denmark and its bearing on the pre-Weichselian occupation of northern Europe by humans

The ecological tolerances of Neandertals, their ability to subsist in the dense forests of full interglacials, and their capacity to colonize northern latitudes are the subject of ongoing debate. The site of Hollerup (northern Denmark) lies at the northern extreme of the Neandertal range. Dated by various techniques to the Eemian interglacial (MIS 5e), this site has yielded the remains of several purportedly butchered fallow deer (Dama dama). Taphonomic reanalysis of the remains from Hollerup and a handful of other Eemian-aged fallow deer skeletons cast doubt on the interpretation that they were humanly modified. We place this revised conclusion into the wider context of human settlement of southern Scandinavia during the Eemian. Other claims of Neandertal presence in the region rest on candidate Middle Paleolithic artifacts, all of which derive from surface contexts. With the fallow deer material removed as a secure indicator of Neandertal settlement of Denmark during the last interglacial, this lithic material must be viewed with renewed skepticism. While ecological and/or topographic factors may have played an important role in preventing Neandertals from penetrating into peninsular Scandinavia, we caution that geological, taphonomic, research-historical, and demographic factors may have significantly distorted our picture of their occupation in this region.     

A strontium isotope analysis on the relationship between ritual tooth ablation and migration among the Jomon people in Japan

Distinct patterns can be discerned in the extensive ritual tooth ablation found among the human skeletal remains of the Late–Final Jomon period (ca. 3200–2800 cal BP) in Japan. Based on comparative observations of sex and grave patterns in the skeletal remains, two major patterns in ritual tooth ablation, termed type 4I and type 2C, have been assigned to locals and immigrants, respectively. In order to test this hypothesis, strontium (Sr) isotope (⁸⁷Sr/⁸⁶Sr) analyses were performed on human skeletal remains from the Yoshigo shell mound in Aichi Prefecture, central Japan. Plants in the surrounding area were also examined to illustrate the geographic ⁸⁷Sr/⁸⁶Sr distribution. The Sr isotopic variation in human tooth enamel (⁸⁷Sr/⁸⁶Sr = 0.70868–0.71028) was greater than that in human bones (⁸⁷Sr/⁸⁶Sr = 0.70871–0.70943). Individuals with higher Sr isotope ratios in their tooth enamel than seawater Sr values of 0.7092 can be identified as immigrants (36% of population). The presence of these isotopically identified immigrants between both type 2C and type 4I individuals does not support the previous hypothesis. The intra-population ⁸⁷Sr/⁸⁶Sr distribution of tooth enamel of type 2C individuals showed a significantly higher mean ratio than that of type 4I individuals, suggesting a higher proportion of immigrants among the former.     

A Late Holocene vertebrate food web from southern Haida Gwaii (Queen Charlotte Islands, British Columbia)

This study presents isotopic data (δ¹³C and δ¹⁵N from bone collagen) for 104 samples representing 29 vertebrate taxa from late pre-contact through to contact era (ca. 2000 – 100 BP) Haida Gwaii (British Columbia, Canada) from a wide variety of mostly marine organisms. The results demonstrate the considerable isotopic variability that characterizes potential prey items in coastal contexts and underscores the need for baseline faunal data to interpret human subsistence practices. Based on these data, a detailed marine food web was constructed, which provides insight into local ecological conditions. We present a simple method for quantifying the trophic level of ancient fauna using δ¹⁵N of bone collagen. Finally, we discuss the implications of this study for the reconstruction of human diet.     

Neandertals,early modern humans,and rodeo riders

In 1995 Berger and Trinkaus (J. Archaeol. Sci. 22, 841–852) proposed that the anatomical distribution of Neandertal trauma, with a predominance of upper body lesions, reflected close-quarter ambush hunting as dictated by the available Middle Paleolithic weaponry (the “Rodeo rider” hypothesis). The necessity for mobility among these Late Pleistocene foragers, as a factor possibly reducing the number of preserved lower limb injuries, was considered as an alternative explanation. The accumulating data on Upper Paleolithic injuries and Middle Paleolithic weaponry, considerations of differential skeletal susceptibility of minor trauma, and evidence of interhuman violence, plus the importance of mobility for Late Pleistocene human existence, suggest that hunting injuries may explain only part of the pattern. The purpose of this note is not to resolve to ultimate factors behind the anatomical distribution of traumatic lesions among the Neandertals (or early modern humans). It is 1) to emphasize that there are multiple probable contributing factors other than close-quarter ambush hunting due to the limitations of Middle Paleolithic weaponry, and 2) to open the discussion to alternative interpretations.     

Neandertal Behavior at the Middle Palaeolithic Site of Abric Romaní, Capellades,Spain

Abstract

This paper synthesizes the work carried out since 1983 at the Middle Palaeolithic site of Abric Romaní, Spain, reconstructing Neandertal behavioral and settlement patterns. The sedimentary features of this site give a high resolution of prehistoric occupational events. Large surface excavation (over 300 sq m) has been used to obtain a detailed picture of intrasite settlement patterns. The reconstruction of activities and their spatial and temporal organization is a key issue in the debate concerning the cultural capabilities of Neandertals versus anatomically modern human beings, and our interpretation integrates geoarchaeology, lithic technology, taphonomy, zooarchaeology, and anthropology. In this way, we explain the degree of diversity within a single Neandertal archaeological site, especially regarding lithic assemblage variability, faunal processing activities, and settlement patterns.     

Hydrogen isotope systematics of hair: archeological and forensic applications

The hydrogen isotope systematics of hair have been explored and several applications demonstrate the type of information that can be recovered from archeological and forensic hair samples. Experiment with modern hair demonstrate that sources of protein-based hydrogen in keratin are food and water. On the basis of a deuterium-enriched drinking study, it was found that 31% of hydrogen in human hair is derived from ingested water. At least 9% of total hydrogen in human hair is isotopically exchangeable with water or water vapor at 25 °C. Nearly complete exchange occurs in a matter of hours. The δD value of body water is approximately 17±10‰ (n=7) more negative than human hair, although there is much scatter in data for modern hair, which is clearly related to variability of the isotopic composition of ingested food and water. Archeological hair samples were analyzed from (i) a 370 year old Incan mummy sacrificed at 5300 m altitude in Argentina (Geoarchaeology 14 (1999) 27) and (ii) from a Woolly Mammoth from Siberia. High spatial resolution data along the length of hair from the mummy indicate seasonal variations in carbon, nitrogen and hydrogen isotope ratios. No hydrogen isotope anomalies are seen in the youngest hair at the base of the scalp, indicating that the individual probably lived at 1600 m, and spent less than a week at high altitudes before death. The mammoth also preserves a seasonal signal, suggesting hair growth rates of 33 cm/year, faster than human hair, but slower than horse tail hair. The limited isotopic range across seasons for the mammoth is consistent with a north-south migration on the order of 1000 km.     

Application of magnetic susceptibility on polished stone tools from Western Hungary and the Eastern part of the Czech Republic (Central Europe)

A non-destructive method – magnetic susceptibility (MS, κ) measurement – was used to characterize about 360 Neolithic, Copper and Bronze age polished stone tools of the Miháldy collection (Laczkó Dezső Museum, Veszprém, Hungary), composed of artefacts from three Neolithic (Moravian Painted Ware) workshops around a source of greenschist at Želešice near Brno (Moravia, Czech Republic) and semifinished artefacts from workshops at Neolithic mines in the Jizerské hory Mts. (Bohemia, Czech Republic); in addition, rock reference samples collected from the possible provenance areas and rock outcrops in the Czech Republic, Slovakia and Hungary were analysed as comparative material with the same method. We elaborated a model experience to control the thickness correction factor (TCF) following Williams-Thorpe et al. (2000).Among various stone raw materials of polished artefacts we found basaltic rocks and greenschists-metabasites as the most common.     

Human and animal diet at Conchopata, Peru: stable isotope evidence for maize agriculture and animal management practices during the Middle Horizon

This paper reports δ¹³C and δ¹⁵N values for human and animal skeletal remains from the Middle Horizon (AD 550–1000) site of Conchopata in the Peruvian highlands. The data indicate that maize was the dietary staple for both humans and the majority of animals at this urban site. Camelids at the site segregated into two groups according to δ¹³C values, reflecting two distinct animal management strategies. Human infants were found to have elevated δ¹⁵N values, reflecting the trophic level effect of breastfeeding. No sex based differences in diet were observed.     

Vote evolution in Spain, 1977–2007: A spatial analysis at the municipal scale

This paper presents a spatial analysis of the results in the general elections of contemporary Spain (1977–2007). By using electoral data at municipal level, we have traced back the evolution of the vote at high spatial resolution, thus providing a more accurate overview of the evolution of modern Spanish politics and offering empirical evidence on the nationalization thesis. We demonstrate that this approach reveals several processes that remain hidden when aggregated data are used, especially the clustering patterns in the smooth transition from dictatorship to full democracy (the so-called Spanish Transition). We found a statistical normalization process of the left/right dynamics with a growing number of balanced municipalities with a similar number of votes for left and right parties. The tendency towards a Gaussian histogram centered on a value of 50% for the left vote is an indication of political normalization after 40 years of dictatorship.     

Radiocarbon Chronology of the Siberian Paleolithic

We have compiled 462 C-14 determinations for 120 Paleolithic and Mesolithic sites from Siberia and the Russian Far East. The Mousterian sites are dated to ca. 46,000–28,500 BP. The Middle–Upper Paleolithic transition dates to ca. 43,300–28,500 BP. Although there are a few earlier sites, most of the Upper Paleolithic sites are dated to the time interval between ca. 34,000 BP and 10,000 BP. The earlier Upper Paleolithic stage is characterized by macroblade technology and is radiocarbon-dated to ca. 34,000–20,000 BP. The earliest microblade technology occurs in the late stage of the Upper Paleolithic, dated to ca. 23,000–20,000 BP, but the majority of microblade sites is dated to ca. 20,000–11,000 BP. The Final Paleolithic (Mesolithic) sites date to ca. 12,000–6000 BP. At ca. 13,000–11,000 BP, the earliest Neolithic appeared in both the Russian Far East (Amur River basin) and the Transbaikal. The Paleolithic–Neolithic transition occurred ca. 13,000–6000 BP.     

Preliminary neutron diffraction study of two fenestrated axes from the ‘Enot Shuni' Bronze Age cemetery (Israel)

Neutron diffraction (ND) analyses of ancient metals show that this method is capable of detecting differences in the inner composition and microstructure of ancient metal objects. Here, ND measurements were conducted on two ‘eye shaped’ axes from the end of the 3rd-beginning of the 2nd millennium BC. The objects were excavated from the ancient cemetery of ‘Enot Shuni' Israel; one is made of bronze and the other of silver. Both artefacts are rare finds, with the silver axe unique in the archaeology of Israel, and therefore had to be analysed locally. For that purpose, a newly assembled diffractometer (KARL) at the IRR-1 of the Nuclear Research Centre (Soreq, Israel) was used. ND measurement on the bronze axe revealed the existence of an α-phase with a range of Cu/Sn ratios (Cu–Sn solid solutions) and some amount of a δ-phase (intermetallic compound of Cu and Sn). The silver axe ND pattern shows the existence of an α-phase (Ag–Cu solid solution) and some amount of copper metal. Our ND data are discussed in comparison with XRF surface measurements and thermal neutron radiography. The results are shedding more light on the in-depth material composition profile, as well as on the objects’ structural and compositional affinities, and help to better understand the production processes and assist in conservation decisions.     

Approaches to Prehistoric Diet Breadth, Demography, and Prey Ranking Systems in Time and Space

Zooarchaeological data on small game use hold much potential for identifying and dating Paleolithic demographic pulses in time and space, such as those associated with modern human origins and the evolution of food-producing economies. Although small animals were important to human diets throughout the Middle, Upper, and Epi-Paleolithic periods in the Mediterranean Basin, the types of small prey emphasized by foragers shifted dramatically over the last 200,000 years. Slow-growing, slow-moving tortoises, and marine mollusks dominate the Middle Paleolithic record of small game exploitation. Later, agile, fast-maturing animals became increasingly important in human diets, first birds in the early Upper Paleolithic, and soon thereafter hares and rabbits. While the findings of this study are consistent with the main premise of Flannery's Broad Spectrum Revolution (BSR) hypothesis (Flannery, K. V. (1969). In Ucko, P. J., and Dimbleby, G. W. (eds.), The Domestication and Exploitation of Plants and Animals, Aldine Publishing Company, Chicago, pp. 73–100), it is now clear that human diet breadth began to expand much earlier than the Pleistocene/Holocene transition. Ranking small prey in terms of work of capture (in the absence of special harvesting tools) proves far more effective in this investigation of human diet breadth than taxonomy-based diversity analyses published previously. Our analyses expose a major shift in human predator–prey dynamics involving small game animals by 50–40 KYA in the Mediterranean Basin, with earliest population growth pulses occurring in the Levant. In a separate application to the Natufian period (13–10 KYA), just prior to the rise of Neolithic societies in the Levant, great intensification is apparent from small game use. This effect is most pronounced at the onset of this short culture period, and is followed by an episode of local depopulation during the Younger Dryas, without further changes in the nature of Natufian hunting adaptations. An essential feature of the diachronic and synchronic approaches outlined here is controlling the potentially conflating effects of spatial (biogeographic) and temporal variation in the faunal data sets.     

ENSO influence on Holocene Aboriginal populations in Queensland, Australia

In the Pacific region, the onset of modern El Niño/Southern Oscillation (ENSO) activity at approximately 5000 years ago may have played a significant role in the development of cultures in the Pacific basin. Within Australia, similar trends in population and resource use have been identified but largely ascribed to cultural changes. To test human responses to changing ENSO activity through the Holocene we analysed a comprehensive suite of 710 radiocarbon ages from archaeological sites in ENSO-sensitive Queensland. We observe a dramatic and sustained increase in landscape activity at inland sites from 4860 ± 15 years ago, statistically indistinguishable from the timing of the onset of modern ENSO activity. Subsequent changes in long-term activity directly impacted on human populations indicating that once established, ENSO maintained a continuous influence on disparate cultures throughout the Pacific basin.     

Extra-territorial interventions in conflict spaces: Explaining the geographies of post-Cold War peacekeeping

The period since the end of the Cold War has presided over a dramatic expansion in the number of multilateral peacekeeping operations (PKOs). Yet individual states have varied significantly in their enthusiasm for peacekeeping and, moreover, demonstrated a greater propensity to participate in operations located in certain countries than others. Our contribution in the present paper is to provide new insights into how geography underpins these spatial variations. Uniquely, we make use of a geographically disaggregated dataset of multilateral PKOs, which allows us to capture various dyadic linkages between sending and receiving countries. Our results confirm previous work indicating that more democratic countries are more likely to participate in PKOs, but extend these findings by showing that countries' commitment to human rights has a similar positive influence. We also show that aspects of spatial proximity (physical distance, same region) and relational proximity (colonial ties) between potential sending and receiving states raise the likelihood of participation. Yet we find that two relational variables widely discussed in the literature as possible correlates of peaceful interactions – bilateral trade and joint membership of intergovernmental organisations (IGOs) – have no statistically discernable influence on countries' involvement in particular PKOs.