Specialized Early Upper Palaeolithic Hunters in Southwestern France?

Paul Mellars has long used cave and rockshelter ungulate faunal assemblages from southwestern France to argue that the early Upper Palaeolithic people of this region focused their hunting on reindeer (Rangifer tarandus), and that such specialized hunting distinguishes the Upper from the Middle Palaeolithic in at least this region. We examine this argument quantitatively, using a sample of 133 Mousterian, Châtelperronian, and Aurignacian ungulate assemblages. We show that only five Aurignacian assemblages, from three sites, stand out in terms of the degree to which their ungulate faunas are dominated by a single taxon. We also show that some Mousterian cave and rock shelter ungulate assemblages are more heavily dominated by large bovids than Aurignacian assemblages are dominated by reindeer, and that Mellars' argument is highly dependent on the exclusion of open sites from the analysis and on the numerical threshold he has selected to indicate hunting specialization.     

Insights into early Middle Palaeolithic tool use and hafting in Western Europe. The functional analysis of level IIa of the early Middle Palaeolithic site of Biache-Saint-Vaast (France)

Results are presented from the functional analysis of a large selection of tools from level IIa of the early Middle Palaeolithic site of Biache-St-Vaast (France). Results indicate that the site represents a hunting stand with a strong focus on hunting and animal processing activities, next to a maintenance component that appears to concern retooling and repair activities in preparation of the hunting episode(s). In spite of the site's age, a large part of the tool assemblage proves to have been used while hafted. Expertise in hafting was even quite elaborate in the sense that it includes tools for which hafting is not essential. It confirms the importance of hafting studies for improving insights into assemblage variability and past human behaviour.     

Site function and the ‘ibex-site phenomenon’: myth or reality?

Summary.  This paper focuses on ibex hunting in Italy during the Upper Palaeolithic. The faunal assemblages from five ibex-dominated sites (Dalmeri, Arene Candide, Fumane, Villabruna and Soman) are examined with the aim of assessing the role of the sites and people's activities in the wider settlement system. These sites will be placed in their wider context of the Italian Upper Palaeolithic so as to explore the nature of variability in ibex exploitation. The importance of these considerations is to examine aspects of specialized hunting, increased intensity of resource exploitation during the end of the Upper Palaeolithic, and methodological considerations of the relationship between faunal remains and the human activities that produced them.     

Selection and Exploitation of Macro‐Vertebrate Resources During the Upper Palaeolithic in Northern Spain. New Evidence from Coímbre Cave (Peñamellera Alta,Asturias)

The exploitation of ungulates in the Cantabrian region during the Upper Palaeolithic is characterized by the appearance of progressively specialized hunting strategies, especially during the Magdalenian. This specialization focused on either Iberian ibex or red deer, depending on environmental or topographic features. Red deer, for instance, was hunted mostly on the plains while ibex and/or chamois was hunted in rocky and mountainous areas. Here we present new zooarchaeological and taphonomic evidence from Coímbre cave (northern Spain), a site located in the rugged region between the Picos de Europa and Sierra del Cuera (Asturias) which has evidence for specialized ibex hunting. We discuss the possible reasons for such a selective hunting pattern. While the predominance of mountain species such as Iberian ibex or chamois in the Magdalenian levels suggests prey selection based on topographic or environmental criteria, the predominance of large bovids in the Gravettian level could imply that other alternatives were available. We also provide evidence of a pattern of rabbit exploitation which is unusual by comparison with other Upper Palaeolithic sites of northern Spain, and which taphonomic evidence suggests was due to human activity.     

Evidence from the Yana Palaeolithic site,Arctic Siberia,yields clues to the riddle of mammoth hunting

It has become commonplace to talk about humans hunting mammoths, and overhunting is thought to have been one of the causes of the mammoth extinction. However, definite evidence of mammoth kills by humans remains surprisingly scarce. Here we show convincing evidence of mammoth hunting in the Siberian Arctic between 29 000 and 27 000 ¹⁴C years BP. Our data set, from the Yana Upper Palaeolithic site (Siberian Arctic), includes the following: fragments of lithic points and ivory shaft embedded in two mammoth scapulae; two identical holes made by projectiles in a mammoth scapula and a pelvic bone; mammoth tongue bones found in the cultural layer far away from the main mammoth bone accumulation, indicating the consumption of fresh mammoth meat; and a narrow mammoth bone size distribution, implying hunting selection based on animal size. The data suggest that Palaeolithic Yana humans hunted mammoths sporadically, presumably when ivory was needed for making tools. Such non-intensive hunting practiced by humans over millennia would not be fatal to a sustainable mammoth population.     

A Self-triggered Device to Catch Elk as Early as the Neolithic: A Study from an Archaeological and Ethnological Point of View

The paper deals with equipment and methods used by Stone Age man in hunting elk in the extensive woodlands of interior Norrland, northern Sweden. The prevailing division into active hunting and trapping is applied. Ethnological source material is used to exemplify various ways of hunting elk. The methods discussed are: (1) active hunting on skis in winter, (2) driving the animal into an enclosure on bare ground, (3) catching in a pitfall, (4) snaring, and (5) trapping by a self-triggered spear/arrow. A rock carving at Na¨mforsen, Ångermanland, northern Sweden, is evidence that a self-triggered device to catch elk with a spear or arrow was in use even in the Neolithic. It is generally known that the elk was a very important prey for Stone Age man in the forests of Norrland. This is evident from figural portrayals and food remains alike. It is easy to suspect other uses for elk, but they are difficult to prove. Occasionally, however, parts of tools made from elk antler and bone have been found.     

Mammalian fauna from Palaeolithic sites in the Upper Yenisei River basin (southern Siberia): Review of the current zooarchaeological evidence

The palaeontological data on mammal remains from two Middle Palaeolithic (ca. 125 000 years ago – 27 200 BP) and 34 (ca. 34 000–12 000 BP) Upper Palaeolithic cultural complexes in the upper part of the Yenisei River basin (southern Siberia) are considered. General features of the faunal assemblages are established. Several issues are discussed, including (a) changes in species composition through time and palaeoenvironmental implications of the zooarchaeological records; (b) patterns of human exploitation (hunting) of mammals; (c) issue of mammoth hunting and (d) possibility of domestication of dog in the late Upper Palaeolithic in the Yenisei River basin. Copyright © 2009 John Wiley & Sons, Ltd.     

Did Modern Human Dispersal Take a Coastal Route into India? New Evidence from Palaeolithic Surveys of Kachchh,Gujarat

The Indian Ocean coastline is argued to have been a critical route of modern human dispersal from Africa, introducing Late Palaeolithic industries into South Asia, but a dearth of evidence has prevented a direct evaluation of this. Kachchh (Gujarat, India), located immediately east of the Indus Delta, is an important setting to appraise the Palaeolithic occupation of the western Indian coastline. Targeted survey of Late Pleistocene sediments there found widespread evidence for occupation of Kachchh during the Late Pleistocene: Middle Palaeolithic and possibly Late Acheulean lithic artifacts. The conspicuous absence of Late Palaeolithic industries in these Late Pleistocene deposits, with their presence only noted in Holocene contexts, does not support the model of modern human expansions into India with these industries via the coastal route. We evaluate these results in the context of current debates regarding Late Pleistocene hominin demography, adaptation, and expansions.     

Hunting with Howiesons Poort segments: pilot experimental study and the functional interpretation of archaeological tools

For many decades the use of backed pieces from the Howiesons Poort, between about 70 ka and 55 ka ago, in South Africa has been a point of discussion. Recently direct evidence has been provided to associate these tools with Middle Stone Age hunting strategies. Yet, whether they were used to tip hunting weapons or as barbs remained an open question. In this paper we introduce a set of pilot experiments designed to test the effectiveness of Howiesons Poort segments, the type fossils of the industry, hafted in four different configurations as tips for hunting weapons. It is shown that the morphological type can be used successfully in this way. We present the results of a macrofracture analysis conducted on the experimental tools and compare these to results obtained from three Howiesons Poort backed tool samples. By correlating experimental outcomes, macrofracture data and the interpretation of micro-residue distribution patterns, we provide some insight into the functional variables that might be associated with Howiesons Poort segments.     

Hard core and cutting edge: experimental manufacture and use of Magdalenian composite projectile tips

The technology of the European Upper Palaeolithic yielded abundant evidence of the use of composite projectile heads, in the form of osseous points on the side of which one or several (micro)lithic elements are attached. Yet, little experimental work has been devoted to testing and assessing the parameters of use of this type of composite tips. In this paper we present a pilot experiment with replicas of Magdalenian composite spear tips, made of an antler point with one or two rows of flint backed bladelets. Two series of replicas were manufactured after the lithic and osseous record of, respectively, the Lower Magdalenian from southwest France (c. 20–18 Ky cal BP) and the Upper Magdalenian of Pincevent in the Paris Basin (c. 15–14 Ky cal BP). The 34 experimental composite heads were hafted to spears that were then shot with a spearthrower at the carcasses of two young deer. The results provide some insight into the performance characteristics of the osseous and lithic components, both in efficiency and durability. Finally, possible improvements of the experimental protocol are discussed, as well as the implications of our results for the understanding of projectile point variability in the Upper Palaeolithic.     

A crossbeam co-ordinate caliper for the morphometric analysis of lithic nuclei: a description,test and empirical examples of application

Over the last four decades, there has been surprisingly little advance in the quantitative morphometric analysis of Palaeolithic stone tools, especially compared to that which has taken place in biological morphometrics over a comparable time frame. In Palaeolithic archaeology's sister discipline of palaeoanthropology, detailed quantitative morphometric, geometric morphometric, and even 3D geometric morphometric analyses are now seen almost as routine. This period of relative methodological stasis may have been influenced by the lack of homologous landmarks on many lithic tools (essential for any comparative analysis), especially core-based technologies of the Lower Palaeolithic. Archaeological field conditions may also prohibit the use of expensive and delicate precision instruments in certain cases. Here we present a crossbeam co-ordinate caliper that – crucially – both geometrically locates and measures distances between morphologically homologous landmarks upon lithic nuclei via a single protocol. Intra- and inter-observer error tests provide evidence that error levels associated with the instrument fall within acceptable ranges. In addition, we present empirical examples of application in the form of a multivariate analysis of 55 discrete morphometric variables, and a 3D geometric morphometric analysis of co-ordinate landmark configurations derived from Pleistocene lithic nuclei (i.e. ‘cores’ sensu lato). We also introduce to lithic studies some techniques for the study of shape variation that have previously been used with success in biological morphometric analyses. We conclude that use of an instrument such as the crossbeam co-ordinate caliper may provide a useful adjunct to traditional techniques of lithic analysis, particularly in developing a quantitative morphometric approach.     

Re–evaluating Subsistence Skills of Levantine Middle and Upper Palaeolithic Hunters: A Comparison of the Faunal Assemblages

It has long been argued that there were qualitative differences in the procurement strategies of the Middle and Upper Palaeolithic, with the hominids of the earlier period being less capable and efficient hunters than those of the subsequent period. This hypothesis is tested here through an examination of faunal assemblages from the two periods in the Levant by comparing measures of species diversity. It is concluded that no significant differences in hunting can be documented and that the hominids of both periods can be characterized as efficient foragers.     

Rabbits as Food at the end of the Upper Palaeolithic at Molí del Salt (Catalonia,Spain)

Many factors have been causally linked to the diversification of hunting during the European Palaeolithic: declining supplies of high‐ranked prey, considerable human demographic growth, reduced residential mobility, larger populations of ubiquitous small mammals and significant technological developments. However, small prey exploitation was not uniform: the European rabbit (Oryctolagus cuniculus ) is the most frequent species in the Upper Palaeolithic archaeological record of the Iberian Peninsula – south and Mediterranean area – and Southern France. This is demonstrated at Molí del Salt, an Upper Palaeolithic site located at Vimbodí (Catalonia, Spain), whose mammal fauna stands out for the predominance of rabbits [91% of minimum number of individuals (n = 136)]. We analysed the faunal remains from one level [A^sup (c. 12 700–13 000 cal BP)] in order to identify the agent responsible for the faunal accumulation, and to reconstruct aspects of procurement and consumption that shed light on Palaeolithic subsistence strategies in the Northeast Iberian Peninsula. Our results indicate that human agency rather than carnivore activity was responsible for the bone accumulation at Molí del Salt. We identified all the stages in the consumption sequence from skinning to ingestion. We argue that the rabbits were mostly harvested during summer or winter or both seasons. Clearly, the European rabbit was a target species for the human groups which lived at Molí del Salt providing meat, and skin. Copyright © 2016 John Wiley & Sons, Ltd.     

On Neanderthal subsistence strategies and land use: A regional focus on the Rhone Valley area in southeastern France

This study of Middle Palaeolithic assemblages from the Rhone Valley, in the South-East of France, increases our understanding of Neanderthal subsistence strategies and modes of territorial organisation by comparing a wide corpus of human occupations in limited chronological and geographical frameworks. The Neanderthal occupation modes may be examined using sites located in a reduced area, linking medium-altitude territories (Massif Central and the Alpine foothills) to the Rhone corridor. Through the combined analysis of the occupation levels of ten sites, all dated to between Marine Isotopic Stage (MIS) 7 and the beginning of MIS 3, we identify three types of occupation durations: (1) type 1 as long-term residential camps, (2) type 2 as short-term regular hunting camps and (3) type 3 as brief stopover camps. We discuss this variability of habitat types according to various parameters: site age, technical behaviour, environmental conditions, and site localisation and occupation seasons. The aim is to discern the underlying motivations behind Neanderthal group mobility. One of the main features of the Rhone Valley area is the great homogeneity of behaviours reflected within the sequences. This homogeneity, linked to the variability of the site occupations, supports the hypothesis of Neanderthal groups anticipating their land use requirements, and furthermore suggests that another type of circulating model was used in this area.     

Experimental evidence for lithic projectile injuries: improving identification of an under-recognised phenomenon

Between the Upper Palaeolithic and the spread of metallurgy stone-tipped projectiles were of great importance both for subsistence and as weapons. Whilst finds of embedded projectile points in human and animal bone are not uncommon, identifications of such wounds in the absence of embedded points are rare. Previous experimentation involving archaic projectiles has not examined the effects of stone-tipped projectiles on bone. This paper presents the results of experiments in which samples of animal bone were impacted with flint-tipped arrows. The results demonstrate that positive identifications can be made, both grossly and microscopically, of bony trauma caused by flint projectiles. In addition, flint projectiles are shown to often leave small embedded fragments, which can also be identified microscopically. These results compare well with archaeological examples of suspected ‘arrow wounds’ and the article demonstrates the practical application of this data in identifying such injuries. By facilitating the recognition of projectile trauma these findings will have significance both for the investigation of hunting strategies and levels of conflict amongst early human societies.     

Demography and the Palaeolithic Archaeological Record

Demographic change has recently re-emerged as a key explanation for socio-cultural changes documented in the prehistoric archaeological record. While the majority of studies of Pleistocene demography have been conducted by geneticists, the archaeological records of the Palaeolithic should not be ignored as a source of data on past population trends. This paper forms both a comprehensive synthesis and the first critical review of current archaeological research into Palaeolithic demography. Within prevailing archaeological frameworks of dual inheritance theory and human behavioural ecology, I review the ways in which demographic change has been used as an explanatory concept within Palaeolithic archaeology. I identify and discuss three main research areas which have benefitted from a demographic approach to socio-cultural change: (1) technological stasis in the Lower Palaeolithic, (2) the Neanderthal-Homo sapiens transition in Europe and (3) the emergence of behavioural modernity. I then address the ways in which palaeodemographic methods have been applied to Palaeolithic datasets, considering both general methodological concerns and the challenges specific to this time period. Finally, I discuss the ability of ethnographic analogy to aid research into Palaeolithic demography.     

Possible evidence of mammoth hunting during the Epigravettian at Yudinovo, Russian Plain

Whether or not mammoth hunting was practised during the Late Palaeolithic has been a controversial issue ever since large accumulations of woolly mammoth bones associated with prehistoric artefacts were discovered more than 100 years ago. Detailed taphonomic and palaeobiological analyses of the mammoth bone complexes from the Epigravettian Yudinovo site in the Russian Plain were carried out. The combination of the homogeneous weathering rate of the mammoth bones, the isolated state of most of the skeletal elements, the restricted spatial range of the carnivore gnawing traces, the breakage pattern of the skulls and long bones, the sex ratio, the small body size of the adult mammoths, the age profile (with an important frequency of prime-aged cows), and the large number of individuals, suggest that the bone complexes at Yudinovo were constructed from body parts and bones that were extracted from freshly killed mammoths and that mammoth hunting was practised at this site during the Epigravettian.     

Assessing the macrofracture method for identifying Stone Age hunting weaponry

Macrofracture analysis is an experimentally derived method used as an initial step in investigating the hunting function of stone artefacts. Diagnostic impact fractures, which can only develop as a result of longitudinal impact, underpin this method. Macrofracture analysis recently gained favour in Middle Stone Age studies, supporting hypotheses for effective hunting during the late Pleistocene in southern Africa. However, the factors affecting diagnostic impact fracture formation and the interpretation of these fracture frequencies are not yet fully understood. This paper outlines a set of experiments designed to test macrofracture formation under human and cattle trampling and knapping conditions. The results show that: (a) macrofractures occur frequently when stone artefacts are trampled by cattle and humans and in knapping debris; (b) diagnostic impact fractures occur on some of the trampled flakes and knapping debris (≤3%), but significantly less often than in previous hunting experiments; (c) when they do occur, they are likely produced by longitudinal forces similar to those experienced during hunting; (d) considering artefact morphology is important during macrofracture analysis; and (e) macrofracture analysis is not a standalone method, but is most useful as part of a multi-analytical approach to functional analysis. These experiments help to establish a significant baseline diagnostic impact fracture frequency for the interpretation of archaeological macrofracture frequencies.     

The Exploitation of Plant Resources by Neanderthals in Amud Cave (Israel): The Evidence from Phytolith Studies

The depositional environments of Amud Cave indicate that phytolith assemblages retrieved from the cave's sediments are an integral part of the Middle Palaeolithic sequence. As such, they provide direct evidence for plant use. The Amud Neanderthals emphasized both wood and grass exploitation. Ligneous parts of trees and shrubs were used mainly for fuel. Herbaceous plants were used for bedding, possibly fuel, and for food. There is clear and repetitive evidence for the exploitation of mature grass panicles, inferred to have been collected for their seeds. These findings suggest that, as with the pattern recently discerned for faunal resources, a broad spectrum of plants has been exploited from at least the end of the Middle Palaeolithic. Phytolith analysis now provides a tool for testing models explaining subsistence and mobility patterns during the Levantine Middle Palaeolithic and for better understanding the role of vegetal resources in shaping these patterns.     

Hafted armatures and multi-component tool design at the Micoquian site of Inden-Altdorf,Germany

Excavation of the Micoquian site Inden-Altdorf (Weisweiler-124) near the former German capital Bonn in western Germany has revealed the first valid open-site habitation features with hut-like structures and associated hearths for the Middle Palaeolithic in Central Europe. It has been dated to the Eemian interglacial (OIS 5e), a warm interglacial between 128 and 115 ka BP. Various wear traces and especially organic residues have been detected on a large number of stone tools using microscopic use-wear analysis of lithics recovered from the site. A multi-level analysis developed through an experimental framework and archaeological study using optical light microscopes, scanning electron microscopes and energy-dispersive X-ray microprobes identified the adhering residues as birch pitch. Birch pitch is the oldest synthetically produced material and was used as an adhesive to attach lithic implements to wooden shafts. While such hafting technology is commonly associated with modern humans in the Upper Palaeolithic, the birch pitch residues found on the Micoquian tools of Inden-Altdorf suggest that hafting technologies and the frequent use of multi-component tools already existed in the Middle Palaeolithic, c. 120 ka BP in central Europe.