Founder effect, drift, and adaptive change in domestic crop use in early Neolithic Europe

We document and quantify a significant reduction in crop diversity in the early central European Neolithic using a large multi-site database of archaeobotantical remains we compiled from published Neolithic sites across southwest Asia and Europe. Two hypotheses are proposed to account for the observed changes: one which claims that the different environmental conditions of central Europe selected for a different set of crop choices and strategies than in use in southeast and Mediterranean Europe; and a null hypothesis that explains the change as a drift process associated with a small founding population that subsequently undergoes rapid expansion. Through an agent-based simulation model, we test the null hypothesis and demonstrate that the drop in diversity exceeds that predicted by a drift process. We conclude by re-evaluating the possible adaptive changes underlying crop use in early Neolithic Europe.     

Neolithic cultures of the Ukraine and adjacent areas and their chronology

Considerable progress has been made in the study of the Ukrainian Neolithic within the last 30–40 years. Some 500 settlements and more than 20 cemeteries are now known and show many features in common with Neolithic cultures elsewhere in Europe. The Ukrainian Neolithic can be divided into two distinct zones: the food producers (both agriculturalists and animal husbandmen) occupied the forest-steppe on the right of the Dnieper and in the western regions of the Ukraine; the hunter-gatherer-fishers occupied areas to the north and east. The early Neolithic lasted from the end of the eighth to the beginning of the sixth millennium B.P., and the food-producing economies seem to have spread from the south and west. The middle Neolithic lasted until the first quarter of the fifth millennium B.P. and overlapped in time with some of the Aeneolithic cultures. The Neo-Aeneolithic period includes only the last three-quarters of the fifth millennium B.P.     

Modelling the effect of Mesolithic populations on the slowdown of the Neolithic transition

The expansion of the Neolithic transition in Europe took place gradually from the Near East across the whole continent. At Northern Europe, observations show a slowdown in the speed of the Neolithic front in comparison to other regions of the continent. It has been suggested that the presence of high population densities of hunter-gatherers at the North could have been the main cause for this slowdown. This proposal has recently been described by a mathematical model that takes into account: (i) the resistance opposed by the Mesolithic populations to the advance of Neolithic populations in their territory, and (ii) a limitation on the population growth dynamics due to the competition for space and resources. But these two effects are not equally responsible for the slowdown of the spread. Indeed, here we show that the limitation on the population growth dynamics seems to have been the main cause of the delay of the expansion of farming in Northern Europe.     

A multi-layered prehistoric settlement structure (tell?) at Niederröblingen,Germany and its implications

At least 15 settlement layers, deposited above each other from the early Neolithic (ca. 5300 cal BC) until Roman Imperial Times (ca. 400 cal AD), have been discovered at Niederröblingen, Germany. Constituting the longest duration of prehistoric settlement in central Europe so far known (ca. 4000 yrs), the site is thought to represent the first multi-layered settlement mound (tell?) north of the Alps. This implicates that the occurrence of settlement mound is not limited to the circum-Mediterranean area. The detailed chronology, based on archaeological, radiocarbon and OSL dating and supporting a Bayesian model, shows one clear occupational gap (early Neolithic) and a possible hiatus (Roman Times). Erosion phases of the settlement mound and the deposition of alluvial sediments (since Roman Imperial Times) in the surroundings resulted in an almost complete disappearance of the mound from the recent scenery. Being difficult to detect, similar sites might be present, undetected, in central Europe. Their discovery and study would enable the precise reconstruction of the settlement history as well as of the prehistoric man–environmental interactions at a high resolution.     

Spatial dimensions increase the effect of cultural drift

The transition from hunter-gathering to agriculture (Neolithic) spread gradually across Europe from the Southeast. A reduction in cultural diversity of crop farming practices has been previously observed by comparing pre-LBK Neolithic sites in Greece and the Balkans (dated about 8500 yr BP) to LBK Neolithic sites in Central Europe (dated about 7000 yr BP). The decrease in crop diversity is statistically significant even when considering only the species less likely to have been subject to smaller productivity due to climatic factors (reductions in growing season, temperature, daylight, etc.). This reduction in cultural diversity has not been explained previously. In this paper we show that spatial drift, which occurred on the front of the advancing wave of pioneer settlements, can explain the observed loss of diversity during the LBK range expansion. Our results suggest that spatial dimensions can have a relevant effect also in other case studies in which cultural drift is important.     

Spatial–temporal distribution and geographic context of Neolithic cultural sites in the Hanjiang River Basin,Southern Shaanxi,China

Understanding how to live successfully within our environment is among the most pressing challenges facing contemporary society. This paper probes the problem based on comparative analysis and discusses the relationship between the spatial–temporal distribution of the Neolithic cultural sites and the geographic context in the Hanjiang River Basin in the south of Shaanxi Province, China. Archaeological studies have identified 175 Neolithic cultural sites in the study area, with a sequence of Laoguantai (¹⁴C age 8–7 ka BP), Yangshao (¹⁴C age 7–5 ka BP) and the late period of the Neolithic Age (¹⁴C age 5–4 ka BP). The total number of archaeological sites, the distribution area and the density all showed an early ascending and later descending trend, but the proportion of the number of archaeological sites in the study area to the corresponding value of the entire Shaanxi Province declined sharply. Spatially, these sites were concentrated on the terraces of the Hanjiang River and its main tributaries with an altitude of 400–800 m. Multiple data were integrated to clarify the critical effects of tectonic and geomorphologic conditions on the distribution of the Neolithic sites. Further comparisons revealed the correlation of Holocene climate change and environmental evolution with the Neolithic cultural succession in the study area that ameliorated conditions to generally promote the development of the primitive culture while degeneration coincided with the culture's transition or interruption. The discussion on the origin of the primitive culture and the temporal–spatial distribution corresponding to the regional culture differentiation sheds light on the complex and dynamic human–nature interaction system during the Neolithic Age, thus emphasising the wider field-based investigation and high-resolution reconstruction works of the palaeoclimate and palaeoenvironment in the future.     

The Origins of the Neolithic Along the Atlantic Coast of Continental Europe: A Survey

The main aim of this work is to compare the processes of transition to the Neolithic along the Atlantic coasts of continental Europe. Archaeological data on the late Mesolithic and the early Neolithic in the best known regions (central and southern Portugal, Cantabrian Spain, Atlantic France, the shores of the North Sea, and southern Scandinavia) are discussed. The transition to the Neolithic in Atlantic Europe can be viewed as a relatively late phenomenon, with several interesting particularities. Among those, we point out the fundamentally indigenous character of the processes; the existence of a long availability phase, in which hunter-gatherer groups maintained contact with neighboring agriculturalists and probably were familiar with farming and animal husbandry without applying them in a systematic way; and the later development of megalithic monumental funerary architecture. Finally, the main hypotheses so far proposed to explain the change are contrasted with the available evidence: those that argue that the change derives from economic disequilibrium, and those that opt for the development of social inequality as the fundamental cause.     

Ancient DNA,a Neolithic legging from the Swiss Alps and the early history of goat

Ancient DNA from a Neolithic legging (1st half of the 3rd millennium BC) found at Lenk, Schnidejoch (2750 m a.sl.) in the Swiss Alps has demonstrated, that modern distribution of genetic variation does not reflect past spatio-temporal signatures. The legging was made from the skin of a domestic goat (Capra hircus), belonging to the caprine haplogroup B1, which is marginal in Europe today, but represents a third highly diverse goat haplogroup entering Europe already in the Neolithic. Population expansion of lineage B therefore happened more than 4500 years ago, but their members were at some point almost completely replaced by goats of today's common A and C haplogroups.     

Mobility and the diversity of Early Neolithic lives: Isotopic evidence from skeletons

Isotopic analyses of tooth enamel from early Neolithic skeletons in southern Germany adds diversity to the picture of the Neolithic transition in central Europe, which has often been described as a wholesale shift in diet and technology. Over the past decade, these isotopic studies have suggested some degree of immigration from nearby indigenous groups, as well as social differences within early Neolithic communities that correlate with immigration patterns. In general, there emerges pattern a pattern of patrilocal kinship that is consistent with independent genetic evidence, and anthropologically consistent with the potential identification of Neolithic ‘nuclear families’; and finally, specialisation of subsistence activities, such as livestock herding and cultivating, probably along hereditary lines.     

Neolithic economy in central Europe

This article presents a survey of Neolithic economy, land use, trade, natural environment, and usage of plant and animal resources in central Europe, 5415–2240 B.C. (4500–1800 bc). Early, Middle A and B and Late Neolithic materials are summarized and compared. The earliest farmers expanded from southern Hungary and adjacent areas into central Europe over a relatively short time period, 100–200 years. They occupied areas only with good soils; thus probably hunters and gatherers continued to exist in many regions of central Europe. There is an increase in population and more upland areas are exploited for farming during the Middle Neolithic A and B periods. By the Middle Neolithic B period, low-level hierarchical or ranked societies appear in some regions of central Europe. The Late Neolithic may represent a modification of the mixed farming strategy observed during the earlier periods. Perhaps the herding of domestic animals became more important.     

Neolithic Transhumance in the Black Forest Mountains,SW Germany

Abstract

The presence of Neolithic people in mountainous areas of Central Europe is a debated topic and there is a lack of systematic fieldwork to explore patterns of land use in upland regions. The results of two survey campaigns in the Black Forest mountains of SW Germany aid our understanding of Neolithic expansion into the uplands. An extensive inventory of flint artifacts dating predominantly from the Late Neolithic provides evidence of summer highland pasturing and leaf foddering.     

Ecological constraints on the first prehistoric farmers in Europe

The Neolithic Revolution, which witnessed the transformation of hunter–gatherer groups into farming communities, is traditionally viewed as the event that allowed human groups to create systems of production that, in the long run, led to present-day societies. Despite the large corpus of research focused on the mechanisms and outcomes of the Neolithic transition, relatively little effort has been devoted to evaluating whether particular production-oriented adaptations could be integrated into a broad range of ecological conditions, and if specific cultural traditions differed ecologically. In order to investigate whether the differences between the adaptations and geographic distributions of three major Early Neolithic archaeological cultures are related to the exploitation of different suites of environmental conditions, we apply genetic algorithm and maximum entropy ecological niche modeling techniques to reconstruct and compare the ecological niches within which three principal Neolithic cultures (Impressed Ware, Cardial Ware, and Linearbandkeramik) spread across Europe between ca. 8000 and 7000 cal yr BP. Results show that these cultures occupied mutually exclusive suites of environmental conditions and, thus, were adapted to distinct and essentially non-overlapping ecological niches. We argue that the historical processes behind the Neolithization of Europe were influenced by environmental factors predisposing occupation of regions most suited to specific cultural adaptations.     

Mobility and Social Change: Understanding the European Neolithic Period after the Archaeogenetic Revolution

This paper discusses and synthesizes the consequences of the archaeogenetic revolution to our understanding of mobility and social change during the Neolithic period in Europe (6500–2000 BC). In spite of major obstacles to a productive integration of archaeological and anthropological knowledge with ancient DNA data, larger changes in the European gene pool are detected and taken as indications for large-scale migrations during two major periods: the Early Neolithic expansion into Europe (6500–4000 BC) and the third millennium BC “steppe migration.” Rather than massive migration events, I argue that both major genetic turnovers are better understood in terms of small-scale mobility and human movement in systems of population circulation, social fission and fusion of communities, and translocal interaction, which together add up to a large-scale signal. At the same time, I argue that both upticks in mobility are initiated by the two most consequential social transformations that took place in Eurasia, namely the emergence of farming, animal husbandry, and sedentary village life during the Neolithic revolution and the emergence of systems of centralized political organization during the process of urbanization and early state formation in southwest Asia.

     

Saharan prehistoric basketry. Archaeological and archaeobotanical analysis of the early-middle Holocene assemblage from Takarkori (Acacus Mts., SW Libya)

The paper presents the large set of basketry and other worked fibre artefacts constituting the perishable artefacts assemblage from the Takarkori rock shelter. This site is located in southwestern Libya, central Sahara. Its well-preserved Holocene stratigraphy testifies to human occupations by foraging groups (Late Acacus culture; ca. 9000–7400 uncal years bp; ca. 8300–6100 BCE) and herders (Pastoral cultures; ca. 7400–4500 uncal years bp; ca. 6400–3000 BCE).     

Historical context and chronology of Bronze Age land enclosure on Dartmoor,UK

The upland of Dartmoor, southwest England, is one of the flagship prehistoric landscapes within Britain owing to the excellent survival of extensive prehistoric coaxial field systems. Archaeological surveys and rescue excavations during the 1970s and 1980s did much to further the understanding of this landscape; however, much remains to be explored, in particular the chronology of enclosure, the nature of the pre-enclosure landscape and the relationship between Bronze Age communities and their environment. Reconsideration of this landscape is important, given the place it holds in our understanding of subdivision of the landscape across northwest Europe during prehistory. This paper presents new palaeoecological data recovered as part of an integrated archaeological and palaeoecological project on northeast Dartmoor. The sequences detailed here include the first dated Neolithic period palaeoenvironmental data from within the prehistoric enclosed land on the moor, providing a longer-term context for enclosure. Neolithic groups are implicated in the first establishment of heathland in the study area at around 3630–3370 cal BC. During the early Bronze Age, reestablishment of hazel scrub in the study area implies reduced use of the upland, although it is not clear whether this is local or indicative of the wider landscape. A combination of pollen and fungal spore data indicates a substantial shift to species-rich grassland with grazing animals at c.1480 cal BC in a phase that lasted 400 years. The later Bronze Age and early Iron Age are characterised by low intensity use of the upland. These data provide new chronological data for land cover change on Dartmoor and whilst they broadly confirm existing models of upland land use in later prehistory, their proximity to the standing archaeology affords a more nuanced interpretation of local change.     

Vegetation history at the multi-period prehistoric complex at Ballynahatty,Co. Down,Northern Ireland

Palaeoecological methods can provide an environmental context for archaeological sites, enabling the nature of past human activity to be explored from an indirect but alternative perspective. Through a palynological study of a small fen peatland located within the catchment of a multi-period prehistoric complex at Ballynahatty, Co. Down, Northern Ireland, we reconstruct the vegetation history of the area during the early prehistoric period. The pollen record reveals tentative evidence for Mesolithic activity in the area at 6410–6220 cal BC, with woodland disturbance identified during the Mesolithic–Neolithic transitional period ca. 4430–3890 cal BC. A more significant impact on the landscape is observed in the Early Neolithic from 3950 to 3700 cal BC, with an opening up of the forest and the establishment of a mixed agricultural economy. This activity precedes and continues to be evident through the Mid-Neolithic during which megalithic tombs and related burial sites were constructed at Ballynahatty. Due to chronological uncertainties and a possible hiatus in peat accumulation in the fen, the contemporary environment of the Ballynahatty timber circle complex (constructed and used ca. 3080–2490 cal BC) and henge (dating to the third millennium cal BC) cannot certainly be established. Nevertheless, the pollen record suggests that the landscape remained open through to the Bronze Age, implying a long continuity of human activity in the area. These findings support the idea that the Ballynahatty prehistoric complex was the product of a gradual and repeated restructuring of the ritual and ceremonial landscape whose significance continued to be recognised throughout the early prehistoric period.     

Networks and Neolithisation: sourcing obsidian from Körtik Tepe (SE Anatolia)

This paper details the use of obsidian sourcing to reconstruct networks of interaction (or ‘communities of practice’) amongst populations of south-eastern Anatolia and the Near East in the context of ‘Neolithisation’ during the late 11th–early 10th millennia BC. EDXRF was used to elementally characterise 120 artefacts of Epi-Palaeolithic – Pre-Pottery Neolithic A date from Körtik Tepe in south-eastern Anatolia. Four eastern Anatolian sources are represented, mainly Bingöl A/B and Nemrut Da?, plus the first evidence for the use of Mu? obsidian. When the source data is integrated with the artefacts' techno-typological attributes it is possible to locate the assemblage within an Upper Tigris tradition (with some interesting local differences), which stands in stark contrast to contemporary practices in northern Mesopotamia and the Levant. These local and regional distinctions support recent views of the Neolithic being much more heterogeneous, with a ‘mosaic’ of community-specific/local traditions of subsistence practices, raw material choices and lithic technologies during the Younger Dryas–Early Holocene.     

Hunter–gatherer foraging ranges, migrations, and travel in the middle Holocene Baikal region of Siberia: Insights from carbon and nitrogen stable isotope signatures

Foraging ranges, migrations, and travel among Middle Holocene hunter–gatherers in the Baikal region of Siberia are examined based on carbon and nitrogen stable isotope signatures obtained from 350 human and 203 faunal bone samples. The human materials represent Early Neolithic (8000–6800 cal BP), Late Neolithic (6000–5000 cal BP), and Early Bronze Age periods (∼5000–4000 cal BP) and come from the following four smaller areas of the broader region: the Angara and upper Lena valleys, Little Sea of Baikal’s northwest coast, and southwest Baikal. Forager diets from each area occupy their own distinct position within the stable isotope spectrum. This suggests that foraging ranges were not as large as expected given the distances involved and the lack of geographic obstacles between the micro-regions. All examined individuals followed a similar subsistence strategy: harvesting game and local fishes, and on Lake Baikal also the seal, and to a more limited extent, plant foods. Although well established in their home areas, exchange networks with the other micro-regions appear asymmetrical both in time and direction: more travel and contacts between some micro-regions and less between others. The Angara valley seems to be the only area with the possibility of a temporal change in the foraging strategy from more fishing during the Early Neolithic to more ungulate hunting during the Late Neolithic–Early Bronze Age. However, the shift in stable isotope values suggesting this change can be viewed also as evidence of climate change affecting primary productivity of the Baikal–Angara freshwater system.     

Production and subsistence strategies at El Zafrín (Chafarinas Islands,Spain): new data for the early Neolithic of North-West Africa

Until a few years ago North-West Africa was ignored in our hypothesis on the neolithization of Western Mediterranean. Moreover, the few excavations focused exclusively on the test of stratigraphies in order to obtain chronological sequences. Recently developed archaeological projects allowed the inclusion of this region in the scientific debate on the origin and evolution of its Neolithic, on the contacts with the Iberian Peninsula through the Strait of Gibraltar, as well as on the documentation of ways of live, habitation structures, and economic strategies. The open area excavation of Zafrín, in the Chafarinas Isles – an archipelago located a short distance from the North African coast – and the full analysis of its lithic industry (technology, typology and use-wear), here presented, represents a novelty on the Neolithic research in North Africa. This in turn will permit us to approach subsistence strategies and the way of life developed by the Neolithic communities of the region.     

Birds and burials at Ajvide (Gotland,Sweden) and Zvejnieki (Latvia) about 8000–3900 BP

Animal bones in human burials may reveal aspects of the relationship between animals and humans. This article describes the roles of birds in mortuary practices and in the ideology of Stone Age northern Europe. Bird bones from two large burial sites, Middle Neolithic Ajvide (Gotland, Sweden) and Mesolithic and Neolithic Zvejnieki (Latvia) are investigated with osteological methods. Beads and pendants were fashioned from the wing bones of waterbirds, and used in the decoration of the body or the burial dress. The jay was found in three Neolithic burials at Zvejnieki, and it may have been a totem animal for the Middle Neolithic people at Zvejnieki, and its wings or feathers were presumably attached to dresses and costumes for the dead. Bird remains in burials at Ajvide, Zvejnieki and some other Stone Age cemeteries may indicate similar features in the way of perceiving birds, especially the possible symbolic roles of waterbirds and wings. The findings are discussed from the perspective of the cosmology of historical hunter–gatherer (and herding) groups in modern Russia.