The craniology and relationships of four species of Bos,: 4. The Relationship between Bos primigenius Boj. and B. taurus L. and its implications for the Phylogeny of the Domestic Breeds

The three previous papers in this series Grigson, 1974, Grigson, 1975, Grigson, 1976 were a study of the basic craniology of the genus Bos using B. taurus as an example. This paper deals with B. primigenius from Holocene deposits in northern Europe, studying the relationship of absolute size, cranial proportions, angles and non-measurable characters to age and sex. It concludes that the two size forms of B. primigenius are merely the expression of sexual dimorphism, that the two forms B. primigenius and B. taurus are so similar that they form a continuum and should be considered to be of the same taxon, and that where B. taurus does differ from B. primigenius the differences can usually be ascribed to breed and can be described as specializations.     

The craniology and relationships of four species of Bos 3. Basic craniology: Bos taurus L. Sagittal profiles and other non-measurable characters

This paper is the third in a series on the craniology of four species of Bos and it is the last of the three papers in which the skulls of British domestic cattle are used in an attempt to establish a basic craniology for the genus. Part 1 (Grigson, 1974) dealt with nomenclature, material, history, age assessment and absolute size. Part 2 (Grigson, 1975) dealt with cranial proportions and angles. This paper deals first with sagittal profiles and then with other non-measurable characters.Sagittal profiles may be rounded or pointed with some intermediate forms. The rounded shape (an infantile character) persists throughout life in some breeds and is unaffected by age, sex or size. The pointed form develops with age in some breeds and is probably more marked in older and larger animals. Seventeen other minor characters are examined for the effects of age, sex, breed and size.     

The craniology and relationships of four species of Bos II. Basic craniology: Bos taurus L. Proportions and angles

This paper is the second of a series on the craniology of four species of Bos. It is a further study of the skulls of British domestic cattle which are used in an attempt to establish a basic craniology for the genus. Part 1 (Grigson, 1974) dealt with nomenclature, material, history, age assessment and absolute size. This paper deals first with cranial proportions and then with cranial angles.The results show that the proportions of the skull depend on its size and, to a lesser extent, on its sex, cows having relatively narrow skulls, with short horncores, but sometimes longer horns than bulls. Cranial angles seem to be independent of size or sex, but are influenced slightly by age, and also, in some cases, strongly by breed.     

The craniology and relationships of four species of bos 1. Basic craniology: Bos taurus L. and its absolute size

This paper is the first of a series on the craniology of four species of Bos. Here (and in the next paper) Bos taurus in Britain is used to establish a basic craniology for the genus. In the initial sections notes are made on the nomenclature, material available, history, and age assessment of B. taurus. The subsequent section deals with the effects of sex, age and breed on absolute size.The results show that Bos taurus is very variable, particularly in the length of the horncore. Sex influences certain dimensions very greatly, particularly those of breadth. Although the horns of cows are often longer than those of bulls of the same breed the horncores may be shorter in cows. In studying the effects of age it can be seen that growth rates are initially very high, slowing down at about three years of age. Tooth row length decreases with age and hence it is a particularly unreliable indicator of size. Obviously breed affects size and it has been possible to divide the skulls studied into size groups which correspond well with known body weights.     

Adult and juvenile aurochs, Bos primigenius Boj. from the maglemosian site of ulkestrup lyng Øst,Denmark

Bones of adult and juvenile aurochs (Bos primigenius Boj.) from the refuse of a Mesolithic settlement, Ulkestrup Lyng Øst (c. 6090 BC), are examined. The problem of estimating the ontogenetic age of the younger animals is dealt with in order to establish the time of the year at which they were killed and thereby the possible season of hunting.The age-determination is mainly based on tooth-wear, which is discussed in its relationship to physiological changes in the stomach, that are correlated with the dietary changes from milk to solid food. The subfossil mandibles are compared with modern material from domestic calves and with other Danish subfossil calves' mandibles.     

Molecular markers for the discrimination of Triticum turgidum L. subsp. dicoccum (Schrank ex Schübl.) Thell. and Triticum timopheevii (Zhuk.) Zhuk. subsp. timopheevii

The persistent uncertainty on the classification of the “new” glume wheat found in Neolithic and Bronze Age sites from Greece and other European settlements might be resolved only through analysis of its ancient DNA. Tools able to discriminate among different Triticum species on the basis of scarce, very damaged DNA, are therefore essential. While current attempts concentrate on DNA fragments sequencing and comparison, in some instances PCR-based selective amplification techniques might offer a cheaper and quicker alternative. The purpose of this research was therefore the identification of species-specific primers, able to distinguish caryopses of Triticum timopheevii subsp. timopheevii from those of Triticum turgidum subsp. dicoccum. Primers and their working conditions were defined and optimized using DNA from modern accessions. The ribosomal primers ITS1 tim and ITS2 tim, and the nuclear primer acetyl-coenzyme A tim clearly discriminated the sequences of Triticum timopheevii from other species. Finally, Neolithic charred wheat grains found in the sites of Sammardenchia (Pozzuolo del Friuli, Udine) and La Marmotta (Lago di Bracciano, Roma), belonging to the “new” wheat type or to emmer, were tested with the three selected primers. However, the results were not conclusive, because the samples analysed were apparently too degraded to yield useful DNA.     

The Quaternary fossil record of the genus Testudo in the Iberian Peninsula. Archaeological implications and diachronic distribution in the western Mediterranean

This paper explores the presence of tortoise remains and its meaning in Palaeolithic archaeological contexts. We focus on two topics: (1) the presence or absence of the genus Testudo from the Early Pleistocene to the Holocene in SW Europe; and (2) the relationship between Testudo and the palaeoclimatic and palaeoenvironmental conditions diachronically. In order to address this objective, we collected published and unpublished data from different sites on the eastern Iberian Peninsula. The results obtained by this study demonstrate that Testudo hermanni was the only tortoise present in this area until recent times and its presence in archaeological sites is almost always related to anthropic activities. Finally, the disappearance of Testudo from archaeological sites in some areas is related to climatic changes and events that occurred during the last stages of the Pleistocene.     

Insights into early Middle Palaeolithic tool use and hafting in Western Europe. The functional analysis of level IIa of the early Middle Palaeolithic site of Biache-Saint-Vaast (France)

Results are presented from the functional analysis of a large selection of tools from level IIa of the early Middle Palaeolithic site of Biache-St-Vaast (France). Results indicate that the site represents a hunting stand with a strong focus on hunting and animal processing activities, next to a maintenance component that appears to concern retooling and repair activities in preparation of the hunting episode(s). In spite of the site's age, a large part of the tool assemblage proves to have been used while hafted. Expertise in hafting was even quite elaborate in the sense that it includes tools for which hafting is not essential. It confirms the importance of hafting studies for improving insights into assemblage variability and past human behaviour.