Size and shape of the Eurasian wild boar (Sus scrofa), with a view to the reconstruction of its Holocene history

Abstract

A large assortment of skulls and skeletons of recent wild boar (Sus scrofa) from across the world has been used to collect tooth and bone measurements that can be compared to those from archaeological specimens. The data provide useful information for a reconstruction of the Holocene history of the species. The evidence collected so far highlights the great variability of the species and provides a baseline to be used for the interpretation of ancient material. It is shown that not only the size, but also the shape of teeth and mandibles can help in highlighting patterns of variability in wild boar from different areas. A number of geographic trends are identified in the variation of S. scrofa across its range, mainly concerning the differentiation of insular forms, and the existence of South–North and West–East clines. Other factors such as hybridisation with domestic stock, feralisation and human-induced movement of animals may also play an important role. A comparison with ancient material emphasises the existence of similarities as well as differences between modern and ancient populations. Although some of the geographic trends identified on the basis of the analysis of modern material seem to date back to early Holocene times, the morphological history of the species appears to be complex, and in more than one area fluctuations in body size seem to have occurred.     

Plant macroremains from the Roman harbour of Pisa (Italy)

Abstract

Results relating to the identification of macroscopic plant remains (leaves, fruits, seeds) discovered during the archaeological excavation of the Roman harbour of Pisa, in Tuscany, Italy are reported. The plant samples include the cultivated species Prunus spp., Corylus, Olea, Vicia, Juglans and the nemoral wild species Salix spp., Quercus spp., Fagus, Ulmus, Alnus. The remains of cultivated plants — fragments of fruits and seeds — were perhaps part of the boats' cargo or stores for the crew's meals. In the case of the wild taxa, the simultaneous presence of fruit and leaves, suggests that the remains recovered came from the forest vegetation bordering the area of excavation and the adjacent higher grounds, thus delineating a new vegetation landscape for the ager pisanus.     

A Pilot Study in Odontochronology for the Pig Domestication Question in North‐Central China

In this paper, we report results of the pilot study in which we apply skeletochronological analysis of teeth to a collection of modern Chinese wild boar (Sus scrofa) of known age and date of death. This is the first time that odontochronology has been applied to suids in China. This work allowed us to establish parameters for subsequent analysis of a limited number of Neolithic‐aged Sus (wild boar and possible early domesticated pig) samples from Henan Province. The goal of the research is to contribute to the ongoing reconstruction of Neolithic dietary, health and economic patterns along the Yellow River. Our results are encouraging in that regular seasonal growth patterning is observed in the northern control sample, which is from the same region as our archaeological investigation. Copyright © 2011 John Wiley & Sons, Ltd.     

A supernumerary permanent mandibular premolar of wild boar (Sus scrofa L.) from the early medieval Novgorod,Russia

The aim of this study was to investigate a unique case of polydonty in wild boar from the early medieval Novgorod (second half of 11th century AD). The only preserved distal part of the right mandibular branch shows a rotated transitional permanent extra premolar between P₃ and P₄. This abnormality has the potential to serve as a territorially‐based indicator of past wild boar populations. Copyright © 2009 John Wiley & Sons, Ltd.     

Subsistence continuity,change, and environmental adaptation at the site of Nugljanska,Istria, Croatia

This paper investigates changes in subsistence strategies at the upland cave site of Nugljanska (Croatia) at the end of the last ice age, during a time of rapid sea level rise and changing environment. We analysed the faunal assemblage from archaeological levels spanning the Pleistocene–Holocene transition (15,000–8000 years BP) and evaluated changes in relative abundance of species, measured species diversity, and compared the representation of terrestrial mammal and marine resources. We found a significant shift in the most abundant prey species exploited (from red deer, Cervus, to wild boar, Sus). There was some correlation between dietary diversification and periods of increased moisture availability and the spread of deciduous forest in the area. Our results suggest that there was a continuing reliance on terrestrial resources throughout time and that changes in dietary patterns were likely due to local environmental change and potentially, changing seasonal mobility strategies, at the Pleistocene–Holocene transition.     

The pectoral fin spines of European catfish Siluris glanis; cultural artifacts or food remains?

The urban archaeological site of Nicopolis ad Istrum in north central Bulgaria, which has multiple periods of use and abandonment, has produced a large and varied ichthyofauna. A number of bones of the European catfish Siluris glanis have been identified and have been interpreted as both food remains and bone artifacts. It was found that a percentage bias in the distribution of S. glanis bones from the Roman period of occupation called for further analysis. The investigation attempts to identify the reason or reasons for the apparent overrepresentation of the pectoral fin spines of this species. Taphonomic factors are considered along with butchery and an actualistic study of butchery on modern S. glanis material. Studies from north Africa are used to illustrate the possible function of the spines. The conclusions drawn are that a high proportion of the pectoral fin spines were brought into the site, possibly as personal artifacts. Their use is open to debate; no clear functional use can be determined from the archaeological material, although it cannot be ruled out that they were used for medicinal purposes or as charms.     

Taphonomy and zooarchaeology of the Upper Palaeolithic cave of Dzudzuana,Republic of Georgia

We present the results of a detailed taphonomic and zooarchaeological study of the faunal remains from the Upper Palaeolithic layers of Dzudzuana Cave, Republic of Georgia. This study presents the first carefully analysed Upper Palaeolithic faunal assemblage from the southern Caucasus and thus serves as a significant point of reference for inter‐regional studies of Upper Palaeolithic subsistence in Eurasia. A series of intra‐site taphonomic comparisons are employed to reconstruct the depositional history of the bone assemblages within the different occupational phases at the site and to investigate subsistence, meat procurement and bone‐processing strategies. Caucasian tur (Capra caucasica), aurochs (Bos primigenius) and steppe bison (Bison priscus) were the major prey species throughout the Upper Palaeolithic. Their frequencies do not change significantly over time, and nor does bone preservation vary by layer. The assemblage is characterised by significant density‐mediated biases, caused by both human bone‐processing behaviours and in situ post‐burial bone attrition. Bone marrow extraction produced large numbers of unidentified bone fragments, many exhibiting green bone fractures. The density and size of bone assemblages and the extent of fragmentation indicate that Dzudzuana Cave was repeatedly occupied by Upper Palaeolithic foragers over many years. Skeletal part representation and butchery marks from all stages of carcass processing suggest that prey occasionally underwent field butchery. Intra‐site taphonomic comparisons highlight uniform patterns of cultural and economic behaviours related to food procurement and processing strategies. Copyright © 2007 John Wiley & Sons, Ltd.     

Vertebrate Animal Remains from Prehistoric and Medieval Settlements in Primorye (Russian Far East)

A review of zooarchaeological research is presented for one of the best-studied areas of the Russian Far East, Primorye (Maritime) Province. The faunal remains, including mammals, birds and fish, were derived from archaeological contexts ranging from the Upper Palaeolithic, ca. 33 000 years ago, to the Middle Ages, twelfth to thirteenth centuries AD . Among the wild species, hoofed animals, wild boar, and bears are the most common. Domesticated animals are represented mostly by pig and dog. At the Pleistocene–Holocene boundary, ca. 10 000–12 000 years ago, some species, such as mammoth, woolly rhinoceros and bison, became extinct. Since the Middle Holocene, ca. 7000 years ago, the faunal complexes became of modern composition. The finding of bones of domesticated animals in the Bronze Age, dated ca. 2800–3200 years ago, allows the correlation of the emergence of livestock in Primorye with cultural influences from northern China. © 1997 by John Wiley & Sons, Ltd. Int. J. Osteoarchaeol., 7 : 172–180 (1997) No. of Figures: 2. No. of Tables: 3. No. of References: 37.     

Wild galliform and gruiform birds (Aves,Galliformes and Gruiformes) in the archaeological record of Bulgaria

A total of 453 bones (294 of galliform and 159 of gruiform birds) from 28 archaeological sites are considered here, 34 per cent of the total ornithoarchaeological wild bird finds collected in Bulgaria so far. They cover a period from the Early Neolithic to the Middle Ages. Eleven species (six galliform and five gruiform), 87.5 and 45.5 per cent, respectively, of the Holocene avifauna of both orders, have been identified. One species, Tetrao tetrix, has totally disappeared and three others, Grus grus, Otis tarda and O. tetrax, now breed beyond the borders of Bulgaria and the Balkans. Some of the sites of the large gamefowl species indicate their wider distribution in the past. Phasianus colchicus is present as early as the Eneolithic period. The wild galliform and gruiform species were important as a source of meat for the local population up to the Middle Ages, in spite of widespread breeding of domestic poultry. © 1997 John Wiley & Sons, Ltd.     

Comparative human and deer (Odocoileus virginianus) taphonomy at the Richards site,Ohio

The Richards site is attributed to the Philo phase of the Fort Ancient tradition of the Ohio Valley area. Human skeletal material from the site shows evidence of peri‐ and post‐mortem taphonomic changes, including cut marks, burning and fracturing. Previous analyses have discussed explanations for these changes, including secondary burial, ritual destruction and cannibalism. Researchers have theorised that, allowing for differences in anatomy among species, humans and animals butchered for the same purpose (consumption) will show similar patterns of taphonomic changes associated with butchery. The human remains at the Richards Site were disposed in general midden pits containing mixed cultural debris and faunal remains. White‐tailed deer (Odocoileus virginianus) constitutes approximately 60% of all the faunal bone, indicating that it was a major food resource. To test a cannibalism explanation, a comparative analysis of human‐induced taphonomy in human and deer skeletal remains was performed, using chi‐square and odds ratio tests. If humans were being used as a food resource, the pattern of butchery seen would mirror that of the deer. The analysis described here compares the patterns of treatment and disposal of human and deer skeletal elements at the Richards site, to test whether both species were used as food resources. Similar types of evidence for human‐induced taphonomic changes, including cutting, chopping, burning and breakage, can be seen in both species. However, results indicate that, in general, human remains show much more evidence of perimortem treatment than do deer remains. In fact, the common odds ratio for perimortem treatment in all bones is 3.25, indicating that a human bone is 3.25 times as likely as a deer bone to be affected by burning, cutting or chopping. This probably indicates that perimortem treatment of humans was greater than that necessary simply for butchering for consumption. Copyright © 2005 John Wiley & Sons, Ltd.     

Possible Wild Boar Management during the Ertebølle Period. A Carbon and Nitrogen Isotope Analysis of Mesolithic Wild Boar from Fannerup F,Denmark

This paper presents a stable isotope and radiocarbon study on a total of 85 samples of wild boar (Sus scrofa), humans (Homo sapiens), dogs (Canis familiaris), red deer (Cervus elaphus) and roe deer (Capreolus capreolus) from four Late Mesolithic sites in Jutland, Denmark. Four of the eight Sus scrofa samples from one site, the shell midden of Fannerup F, show markedly enriched carbon and nitrogen isotope values, indicating a dietary intake of a substantial amount of marine food. In contrast to standard interpretations of Late Mesolithic animal economy, we suggest that the enriched values of Sus scrofa may be an indication of management by Ertebølle groups in the area that facilitated access to substantial amounts of marine foods for these wild boars compared to contemporaneous conspecifics. The ¹⁴C dates of the Sus scrofa range from 5290 to 4335 cal BC, suggesting that the management of Sus scrofa developed independently of contact with Neolithic societies. Although the sample size remains small, the interpretation of Late Mesolithic animal management adds to the growing evidence for political and economic complexity in the Ertebølle culture.     

Combining Odontochronology,Tooth Wear Assessment,and Linear Enamel Hypoplasia (LEH) Recording to Assess Pig Domestication in Neolithic Henan,China

Mandibles of Sus scrofa (wild boar/pig) from ritual pits H160 and H208 of Longshangang, a Late Yangshao Neolithic site in Xichuan County, Henan, were analyzed for evidence of domestication. Three methods of dental analysis were applied: odontochronology, tooth wear assessment, and linear enamel hypoplasia (LEH) recording, which provide age at death, season of kill, and season of birth data. We investigate whether: (i) the LEH height frequencies on the second molars of the mandibles correspond with the possibility of double farrowing and (2) double farrowing is supported by the season of and age at slaughter data. If so, these data suggest a substantial degree of human management of suids at the site. Copyright © 2014 John Wiley & Sons, Ltd.     

An Experimental Approach to the Disaggregation of Samples from Peat Deposits

Abstract

Excavations carried out at Dudley Castle recovered an assemblage of animal bones dominated by horse from the vaulted cellar beneath the Great Hall. The deposit dates to c. 1710 and appears to have been the result of partial butchery of several aged horses. The possible interpretations are discussed and it seems likely that the deposit represents the leftover waste of a knacker.     

Molluscan Shell Knives and Experimental Cut-Marks on Bones

Abstract

Experimentation demonstrates that the retouched edges of molluscan shells can be used effectively as butchery knives in the absence of lithic raw materials and leave striations on bone surfaces that may be indistinguishable from cut-marks made by stone knives. The potential of such non-lithic cutting tools suggests one new possible category of early artifact, and may explain the presence of cut-marks on fossil bones in paleoenvironments where stone artifacts were absent or rare.     

Quantitative micromorphological analyses of cut marks produced by ancient and modern handaxes

In this study, we analyse the three-dimensional micromorphology of cut marks on fossil mammal remains from a ∼0.5 million year old Acheulean butchery site at Boxgrove (West Sussex, southern England), and make comparisons with cut marks inflicted during the experimental butchery of a roe deer (Capreolus caproelus) using a replica handaxe. Morphological attributes of the cut marks were measured using an Alicona imaging microscope, a novel optical technique that generates three-dimensional virtual reconstructions of surface features. The study shows that high-resolution measurements of cut marks can shed light on aspects of butchery techniques, tool use and the behavioural repertoire of Lower Palaeolithic hominins. Differences between the experimental cut marks and those on the Boxgrove large mammal bones suggest variation in the angle of the cuts and greater forces used in the butchery of the larger (rhinoceros-sized) carcasses at Boxgrove. Tool-edge characteristics may account for some of these differences, but the greater robusticity of the Boxgrove hominins (attributed to Homo heidelbergensis) may be a factor in the greater forces indicated by some of the cut marks on the Boxgrove specimens.     

First Introduction and Early Exploitation of the Persian Fallow Deer on Cyprus (8000–6000 cal. bc)

The Persian fallow deer (Dama dama mesopotamica) is currently a threatened species. However, it played an important role in many Late Glacial and Early Holocene human societies in the Near and Middle East. This is especially true of the island of Cyprus, where it was introduced at the beginning of the Neolithic and held a predominant place in human subsistence throughout Cypriot prehistory until the Bronze Age. The earliest levels of the extensive Cypriot Pre‐Pottery Neolithic B site of Shillourokambos, occupied between 8400 and 7000 cal. bc , provided 3036 identified remains of this deer. It was possible to measure or determine the age‐at‐death for 1361 and 1444 remains, respectively. Analyses allow for discussions on when the fallow deer was introduced to the island of Cyprus, its origin and how populations were managed. These studies also lead to the reconstruction of acquisition and butchery techniques, as well as culinary practices, and the morphological evolution of males and females throughout time. The Persian fallow deer was introduced to Cyprus later than suids, dogs, cats, goats and cattle, and at nearly the same time as sheep, towards ca 8000 cal. bc . Despite the absence of any skeletal changes, this introduction may reflect an attempt to domesticate the fallow deer on the nearby continental mainland. However, after being introduced to the island, deer appear to have been released into the wild and hunted. Copyright © 2015 John Wiley & Sons, Ltd.     

Shape Change and Variation in the Cranial Morphology of Wild Canids (Canis lupus,Canis latrans,Canis rufus) Compared to Domestic Dogs (Canis familiaris) Using Geometric Morphometrics

Wild canid populations exhibit different anatomical morphologies compared to domesticated dogs in North America. This is particularly important concerning archaeological sites, which may contain early domesticated species, for the proper identification of osteological remains. Previous studies have indicated domestic dogs exhibit a shorter rostrum accompanied by a crowded tooth row; however, none describe the overall complexity of these changes. Consequently, using a landmark‐based geometric morphometric analysis, cranial morphological characteristics were examined in North American wild canids: the gray wolf (Canis lupus), coyote (Canis latrans), red wolf (Canis rufus), and the domestic dog (Canis familiaris). The shape and size of the cranium in lateral and ventral views were compared between the three wild species to the group of domesticated dogs. Wild canids clustered separately from the domestic group in all statistical analyses. Results indicate an expansion of the orbital region, a compression of the rostrum, and an overall warping in the shape and orientation of the skull. In domestic species, there is also a downward shift in the frontal portion of the skull accompanied by the braincase assuming a more upward position. This technique successfully depicted how slight changes in isolated areas of the cranium can have an impact on the overall shape and morphology of the skull. We presume these changes in cranial anatomy reflect the recent selective pressures domestic dogs have undergone since diverging from their wild ancestors. Copyright © 2012 John Wiley & Sons, Ltd.     

Coleoptera from Late Medieval Smoke-Blackened Thatch (SBT): their Archaeological Implications

Abstract

Distinctive assemblages of Coleoptera (beetles) recovered from seven samples of smoke-blackened thatch (SBT)from a range of late medieval roofs in the south of England are listed. The age, ecology and possible origins of this fauna are discussed. Although its precise origin is known, it is suggested that its occurrence is not limited to thatch, and so cannot be used as a 'finger print' for thatch per se. A find of Sitophilus granarius, the 'granary weevil', suggests that thatch may represent another route by which this species has been incorporated into the archaeological record.     

Rewilding – A New Paradigm for Nature Conservation in Scotland?

Abstract

Rewilding is a strategy for the conservation of complete, self-sustaining ecosystems, primarily involving the protection and, where necessary, reintroduction, of populations of keystone species in large, connected reserve networks. A potential method of preserving ecosystem functions and biodiversity, it is now receiving a great deal of practical and political attention, particularly in North America. In Scotland, where many native species have been extirpated in the relatively recent past, rewilding has clear relevance and may provide an overarching set of objectives for current programmes of native woodland restoration and species reintroductions. Nevertheless, rewilding is not widely used as a term or strategy in Scottish conservation. This review considers the development of the concept and its possible application in Scotland, and identifies substantial scope for rewilding, in terms of the restoration and protection of large areas of wild land, and of the reintroduction of native species which have been driven to extinction by human activity. As the environmental, social and economic benefits which are likely to result from a programme of rewilding in Scotland outweigh the potential drawbacks, the adoption of rewilding is recommended as one aim of environmental policy.     

The archaeobotany of Indian pulses: identification,processing and evidence for cultivation

Abstract

Pulses are a significant component of traditional subsistence in South Asia. Reliable identification criteria for identifying these from archaeological seed remains are reviewed. The botanical evidence relating to the wild progenitors and their distribution, especially of Indian natives (Macrotyloma uniflorum, Vigna radiata, Vigna mungo) is summarised, including new evidence from primary botanical research. The problem of seed size increase in pulses is reviewed through a focused study on Vigna spp., in which it is shown that seed enlargement is delayed by 1–2,000 years after initial cultivation. The taphonomy of archaeological pulses is considered in the context of crop-processing of pulses, in which an important distinction can be drawn between free-threshing and pod-threshing types. The total archaeobotanical record for pulses in South Asia (India and Pakistan) is summarised and key regional differences are highlighted.