Avian Remains from Khonkho Wankane,an Urban and Ceremonial Center in the Southern Lake Titicaca Altiplano

The urban and ceremonial center Khonkho Wankane flourished in the southern Lake Titicaca Altiplano region during the Late Formative prior to the rise of Tiwanaku as a dominant urban center. This site yielded an avifaunal number of identified specimens (NISP) = 631, with a minimum of 26 taxa represented among 539 skeletal elements or fragments. A total of 92 egg shell fragments also were recovered. The avifauna inhabited two major ecological zones: Lake Titicaca, its margins/wetlands and other water margins; and the dry Altiplano grassland (or puna). Lake/wetland taxa include Phoenicopterus chilensis (Chilean flamingo), Nycticorax nycticorax (night heron) and multiple taxa in the Order Anseriformes (ducks, geese, etc.) including Chloephaga melanoptera (Andean goose), Anas georgica (yellow‐billed pintail), A. flavirostris (speckled teal) and Oxyura jamaicensis (Andean ruddyduck). Also present are Fulica ardesiaca (Andean coot), Gallinago andina (puna snipe), Himantopus mexicanus (black‐necked stilt) and Charadrius alticola (puna plover). Puna taxa include Nothoprocta cf. ornata (ornate tinamou), Nothura cf. darwinii (Darwin's nothura), Metropelia sp. (ground doves), Athene cunicularia (burrowing owl), and multiple Passeriformes (songbirds, etc.). Taxa with a cosmopolitan distribution include Falco cf. femoralis (aplomado falcon) and Tyto alba (barn owl). Taphonomic analysis indicates that the avifauna were used in ceremonial contexts, including multiple Falco cf. femoralis burials, and for dietary and possibly tool‐making purposes. Other sources of introduction into the site deposits likely include natural mortality of taxa directly inhabiting the site area. These findings are compared to the avifauna recovered from the Formative site of Chiripa along the Lake Titicaca margin, which included a much higher proportion of lake bird fauna. At Khonkho Wankane, the importance of wetland avifauna may have been enhanced by the alteration of the local environment to include qochas, or artificial reservoirs/ponds, and continued despite reliance upon multiple domesticated plant and mammal species. Copyright © 2011 John Wiley & Sons, Ltd.     

Herpetofaunal Remains from Khonkho Wankane,an Urban and Ceremonial Centre in the Southern Lake Titicaca Altiplano: Unique Behavioural Correlates to Osseous Taphonomy

This research reports the herpetological (reptile and amphibian) remains from Khonkho Wankane, an urban and ceremonial centre in the southern Lake Titicaca region of the Bolivian Altiplano, which fluoresced during the Late Formative period. The total n = 1710, including a minimum of six taxa and representing a significant portion of the relatively depauperate modern herpetofauna recorded for the Lake Titicaca area. A lizard of the genus Liolaemus (Tropiduridae) was identified. Amphibians are represented by Andean toad Rhinella spinulosa (Bufonidae) and by at least four taxa of frogs: [cf. Gastrotheca marsupiata (Hylidae) and Pleurodema marmorata or P. cinerea and Telmatobius spp. (Leptodactylidae)]. The latter genus does not include members of the giant variants (T. culeus) that dwell in Lake Titicaca. The question of how these remains were incorporated into the site deposits is examined taphonomically. No indicators of human or small carnivore predation were detected. Significant spatial clustering of amphibian remains was detected, with two clusters contributing n = 1562 or 92.4% of the amphibian total (n = 1690). These concentrations also are characterised by a depressed ratio of cranial versus postcranial elements, which cannot be explained by recovery methods. These concentrations of herpetofaunal remains likely represent in‐burrow deaths of species sheltering for thermoregulatory benefits away from this harsh Altiplano environment, with the lowered recovery of cranial elements hypothesised to have resulted from a lack of ossification. The bulk of herpetofaunal remains recovered therefore likely were recent intrusions into the site deposits and do not represent the residue of human exploitation during the occupation of Khonkho Wankane. Copyright © 2011 John Wiley & Sons, Ltd.     

Hunting,Herding, and Caravanning: Osteometric Identifications of Camelid Morphotypes at Khonkho Wankane,Bolivia

Khonkho Wankane is a ceremonial center located in the southern Lake Titicaca Basin, Bolivia. During the Late Formative period (AD 1–500), its residents practiced agropastoral lifeways and participated in the rise of the state at Tiwanaku. Like at many Andean sites, bones from the family Camelidae are the most abundant large mammal in domestic contexts. Identifying camelid morphotypes represented by these bones carries far‐reaching implications for understanding past hunting, herding, and caravanning practices, and their roles in larger social and economic webs. Identifications were based on a locally focused reference collection, including llamas (Lama glama) from the immediate vicinity of the site, as well as Andean guanacos (Lama guanicoe), a much smaller morphotype than the Patagonian guanacos used in many osteometric studies. Multivariate statistical analyses and incisor morphology identified all four camelid. Different analyses suggest that the crux of osteometry lies in the reference collection, not the statistical test. An additional, very large morphotype likely corresponds to a castrated llama, the preferred cargo animal among modern drovers. The presence of these animals is interpreted as evidence that groups hunted vicuña (Vicugna vicugna) and guanaco, which are not currently present around the site, herded llamas and alpacas (Vicugna pacos), and perhaps organized caravans with castrated llamas. Copyright © 2013 John Wiley & Sons, Ltd.     

Wombats (Vombatidae: Marsupialia) from the Pliocene Chinchilla Sand,southeast Queensland,Australia

Louys, J., 23.3.2015. Wombats (Vombatidae: Marsupialia) from the Pliocene Chinchilla Sand, southeast Queensland, Australia. Alcheringa 39, XXX–XXX. ISSN 0311-5518

The Chinchilla Local Fauna is one of the richest Pliocene vertebrate fossil assemblages in Australia. However, Vombatidae material preserved in the Chinchilla Sand is very poorly known, and no systematic examination of the wombats from Chinchilla has been conducted. Here I review the cranio-dental and mandibular wombat remains derived from Chinchilla. This material includes both adults and pouch-young specimens. At least five species of wombats are preserved in the fluviatile Chinchilla deposits, although a lack of stratigraphically controlled excavations makes it impossible to determine whether all five species were sympatric. Several wombat taxa are revised: Sedophascolomys gen. nov. is formally erected to replace the invalid ‘Phascolomys’; Vombatus mitchelli (Owen) is recognized as a species distinct from Vombatus ursinus (Shaw), and is recorded for the first time from Chinchilla. In addition to Vombatus mitchelli, the Chinchilla Sand also preserves evidence of Phascolonus gigas, Ramsayia magna, Ramsayia lemleyi and Sedophascolomys medius.

Julien Louys [julien.louys@anu.edu.au], Department of Archaeology and Natural History, School of Culture, History, and Languages, ANU College of Asia and the Pacific, The Australian National University, Canberra, ACT 2601, Australia.     

MOLLUSC SHELL SIZES IN ARCHAEOLOGICAL CONTEXTS IN NORTHERN SPAIN (13 200 TO 2600 CAL BC): NEW DATA FROM LA GARMA A AND LOS GITANOS (CANTABRIA)

Shellfish metrical data are a source of information about the exploitation of marine resources in the past. In this study, we propose a methodological approach based on the size structures of different rocky intertidal gastropod species. Three limpet species (Patella vulgata, Patella intermedia and Patella ulyssiponensis) and the toothed topshell Osilinus lineatus are studied from two sites in Cantabrian Spain: La Garma A and Los Gitanos caves over a period of 10 000 years, covering the Pleistocene–Holocene transition. Data are also supplied about a further sea snail species, the periwinkle Littorina littorea (Upper Magdalenian). A reduction in size can be seen, between the upper Magdalenian and the late Neolithic, in the case of the first four species. The explanation for this decline is probably related to the climate change that occurred in the transition between the Late Pleistocene and the Holocene, but it is possible that human impact might also have influenced shell sizes in the Mesolithic and Neolithic.     

Chonetoidea (Brachiopoda) from the Lopingian (Late Permian) of South China

Ten species of the superfamily Chonetoidea from the Lopingian (Late Permian) of South China are described or revised. A review of all recorded Chonetoidea species from the Lopingian (Late Permian) of South China indicates that some 22 species of five genera can be recognised. Species of Tethyochonetes and Neochonetes are characteristic in the lithofacies dominated by mudstone, siltstone or siliceous rocks in the Lopingian and some argillaceous limestone and clay rock facies near the Permian-Triassic boundary. New taxa are Neochoneles (Zhongyingia) subgen. nov., Neochonetes (Huangichonetes) subgen. nov. and Tethyochonetes flatus sp. nov.     

Isotopic Evidence for Camelid Husbandry During the Formative Period at the Pacopampa Site,Peru

ABSTRACT

Camelids were domesticated in the Andean highlands, such as in the puna habitat, and dispersed into lowland areas and the northern Central Andes. As camelids domesticated in a particular region would have had a greater economic benefit than visiting- or hunted wild camelids, it is important to reconstruct the dispersal of camelid husbandry from its initial site throughout the ancient Andean civilisation. We carried out multi-isotope analyses of animal remains recovered from the Pacopampa site to investigate the nature of camelid pastoralism and utilisation. Strontium and oxygen isotope ratios from tooth enamel suggested that camelids in the early Late Formative Period (800–500 BC) were born near the site and remained in the same habitat for up to three years. Although corresponding data for the Middle Formative Period (1200–800 BC) were not available, carbon and nitrogen isotope ratios were statistically different from those of the Late Formative Period, supporting the possibility that the camelids inhabited the highland plateau like puna. It is inferred that in the northern highlands camelids were initially rare and regarded as either tribute or ritual animals, or they were used as pack animals. Camelid husbandry using maize as fodder began during the Late Formative Period at Pacopampa.     

Late Triassic (Carnian) corals from Timor-Leste (East Timor): their identity,setting, and biogeography

Four scleractinian coral taxa are described from limestones within a sandstone-shale séquence correlated with the Late Triassic Babulu Formation, Manatuto township, on the northern coast of Timor-Leste (East Timor). The coral fauna consists of three phaceloid taxa, Paravolzeia tìmorìca gen. et sp. nov., Craspedophyll ramosa sp. nov., Margarosmilia confluens (Münster), and a generically indeterminate solitary taxon attributed to the family Margarophylliidae. Ali four corals are related at various taxonomie levels to Carnian faunas from the Dolomites of northern Italy. Previously, only Norian coral faunas were known from the Triassic of Timor. The fauna exhibits both similarities to and differences from Carnian faunas of the Dolomites and helps confirm palaeogeographic affinities with the western Tethys, although during Late Triassic time Timor lay in the distant southeastern portai of the Tethys. Despite isolation from the western Tethys, the presence of two species foundalso in the Dolomites indicates that larvai dispersai occurred between the two areas.     

A new fossil genus of forest wallaby (Marsupialia,Macropodinae) and a review of Protemnodon from eastern Australia and New Guinea

A new macropodine genus and species, Silvaroo bila, is described from the Pliocene Chinchilla Sand of Queensland. The generic concept of Protemnodon is reviewed, and it is concluded that two Pliocene species previously placed in that genus (bandharr and buloloensis) belong in Silvaroo. Species of Silvaroo resemble the modern forest wallabies of Papua New Guinea (species of Dorcopsis and Dorcopsulus) and also bear close phenetic resemblance to the late Miocene Dorcopsoides fossilis. Forest wallabies are not known from mainland Australia after the middle Pliocene, but occur in the late Pliocene and Pleistocene of New Guinea.     

A new Changhsingian (Late Permian) Rugosochonetidae (Brachiopoda) fauna from the Zhongzhai section,southwestern Guizhou Province,South China

Zhang, Y., He, W.-H., Shi, G.R. & Zhang, K.-X., 2013. A new Changhsingian (Late Permian) Rugosochonetidae (Brachiopoda) fauna from the Zhongzhai section, southwestern Guizhou Province, South China. Alcheringa 37, 221–245. ISSN 0311-5518.

This paper describes 20 species (including three undetermined species) of Rugosochonetidae (Brachiopoda) in an upper offshore fauna from the Permian–Triassic Boundary Zhongzhai section, southwestern Guizhou Province, South China. New taxa are Tethyochonetes sheni, Tethyochonetes cheni, Neochonetes (Huangichonetes) archboldi, Neochonetes (Sommeriella) waterhousei, Neochonetes (Sommeriella) rectangularis and Neochonetes semicircularis.

Yang Zhang [zyan@deakin.edu.au] and G.R. Shi [guang.shi@deakin.edu.au] (corresponding author), School of Life and Environmental Sciences, Deakin University, Melbourne Burwood Campus, 221 Burwood Highway, Burwood, Victoria 3125, Australia; Weihong He [whzhang@cug.edu.cn] (corresponding author) and Kexin Zhang [kx_zhang@cug.edu.cn], State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, 388 Lumo Road, Hongshan, Wuhan 430074, PR China. Received 8.6.2012; revised 19.9.2012; accepted 7.10.2012.     

Significance of Claraia from the Late Permian of South Guizhou,China

Recently collectée material of two Claraia taxa, Claraia zhiyunica Yang et al, 2001 and Claraia sp. nov. from the Late Permian of South China, are described. Late Permian Claraia species are compared with those from the Early Triassic, and the survival of Claraia across the mass extinction period across the Permian- Triassic boundary (PTB) is discussed.     

Geochemical and Petrographic Analysis of Late Bronze Age Cypriot Ceramics (White Slip I and II and Monochrome) from Tell Atchana/Alalakh (Hatay) in the Amuq Valley

White Slip ware, both White Slip I and II, and Monochrome ware are Middle to Late Bronze Age Cypriot pottery types found across a large area of the Eastern Mediterranean region. A vast quantity of these wares has also been uncovered in Tell Atchana/ancient Alalakh in Hatay in southern Anatolia. We analysed a total of 56 White Slip (n = 36) and Monochrome potsherds (n = 20) from Tell Atchana using XRF, ICP–MS and petrographic thin‐section methods. The main aim of the study was to explore the compositional characteristics of the wares and to determine whether they are local imitations of the Cypriot White Slip and Monochrome wares or represent Cypriot exports to this region. The analytical results proved that White Slip I and II were produced from raw clay of mafic and ultramafic source rocks exposed in the Troodos Massif, available in the Limassol area of southern Cyprus and traded to Tell Atchana. Examples of Monochrome ware excavated in Tell Atchana were also imported to the region, most probably from east/north‐east Cyprus. These results demonstrate a close trading connection between Tell Atchana/Alalakh and southern Cyprus during the Middle to Late Bronze Age.     

Late Oligocene to Early Miocene planktic Foraminiferida from Ashmore Reef No. 1 Well,northwest Australia

Forty-eight species and subspecies, belonging to 13 genera of planktic foraminiferids have been identified from the ‘Cartier Beds’ and unnamed calcarenite in Ashmore Reef No. 1 Well. The assemblages indicate that the units correlate with planktic Zones N.3/4 to N.6–7 (Late Oligocene to Early Miocene). One new species, Subbotina cartieri, is described.     

Late Mississippian (Early Carboniferous) brachiopods from the western Daba Mountains,central China

Qiao L. & Shen S.Z., September 2012. Late Mississippian (Early Carboniferous) brachiopods from the western Daba Mountains, central China. Alcheringa 36, 287–309. ISSN 0311–5518.

Fifteen brachiopod species in 12 genera are described for the first time from four intervals in the middle and upper parts of the Zhanpo Formation at the Huoyanxi section near Zhenba in the western Daba Mountains, southern Shaanxi, central China. The Zhenba brachiopod fauna is dominated by diverse and abundant species of Productida, together with some species of Athyridida, Orthida, Orthotetida and Spiriferida. It ranges from late Viséan to Serpukhovian in age based on the presence of Gigantoproductus species in association with diagnostic foraminifera and conodonts. This fauna generally shows palaeobiogeographical links with the palaeoequatorial realm, including Western Europe, the Moscow Basin, the Ural Mountains, Japan, eastern Tibet and South and North China. Its closest palaeobiogeographical affinity is with South China assemblages rather than those of North and Northwest China, therefore, indicating that the Zhenba area was palaeogeographically close to the South China Block and relatively far from the blocks in Northwest China (e.g., the Qilian Mountains and Qaidam Basin, Kunlun Mountains, Tarim Basin and Tianshan Mountains) during the late Viséan–Serpukhovian.     

Late Devonian (Famennian) brachiopod Yunnanella from the Junggar Basin,northern Xinjiang,China

The Late Devonian (Famennian) brachiopod Yunnanella is reported from the western Junggar Basin, northern Xinjiang, on the basis of Yunnanella hanburii (Davidson) and Yunnanella sp.     

Late Pleistocene Crocuta crocuta spelaea (Goldfuss, 1823) clans as prezewalski horse hunters and woolly rhinoceros scavengers at the open air commuting den and contemporary Neanderthal camp site Westeregeln (central Germany)

Late Pleistocene Ice Age Crocuta crocuta spelaea ( Goldfuss, 1823) hyenas from the open-air gypsum karst site Westeregeln (Saxony-Anhalt, central Germany) is dated into the early to middle Late Pleistocene. Hyena clans apparently used the karst for food storage and as “commuting den”, where typical high amounts (15% of the NISP) of hyena remains appear, also faecal pellets in concentrations for den marking purposes. Additionally small carnivores Meles, Vulpes and Mustela appear to have used some cavities as dens. Several hundreds of lowland “mammoth steppe fauna” bones (NISP = 572) must have been accumulated primarily by hyenas, and not by Neanderthals at the contemporary hyena/human camp site. Abundant caballoid horse remains of “E. germanicus Nehring, (1884)” are revised by the holotype and original material to the small E. c. przewalskii horse. Woolly rhinoceros Coelodonta antiquitatis remains are also abundant, and were left in several cases with typical hyena scavenging damages. Rangifer tarandus (11%) is mainly represented by numerous fragments of shed female antlers that were apparently gathered by humans, and antler bases from male animals that were collected and chewed in few cases (only large male antlers) by hyenas. The large quantities of small reindeer antlers must have been the result of collection by humans; their stratigraphic context is unclear but such large quantities most probably resulted from schamanic activities. The hyena site overlaps with a Middle Palaeolithic Neanderthal camp, as well as possibly with a later human Magdalénian site.     

Phylogenetic analysis of a new fossil Notonectidae (Heteroptera: Nepomorpha) from the Late Jurassic of China

Zhang W.T., Yao Y.Z. & Ren D., June 2012. Phylogenetic analysis of a new fossil Notonectidae (Heteroptera: Nepomorpha) from the Late Jurassic of China. Alcheringa, 239–250. ISSN 0311-5518.

A new fossil species Notonecta vetula sp. nov. is described and illustrated using nymph and adult fossil specimens collected from the Upper Jurassic Chijinqiao Formation, Yumen City, Gansu Province, China. A phylogenetic analysis, based on a combination of fossil and extant backswimmers, was conducted to confirm the position of the new fossil within the Notonectidae.

Wei-ting Zhang [zhangweitinghao@163.com], Yun-zhi Yao* [yaoyz100@gmail.com] and Dong Ren [rendong@mail.cnu.edu.cn], Key Lab of Insect Evolution and Environmental Changes, Capital Normal University, Beijing 100048, PR China; *Corresponding author; also affiliated with: State Key Laboratory of Palaeobiology and Stratigraphy (Nanjing Institute of Geology and Palaeontology, CAS), Nanjing 210008, PR China. Received 13.7.2011; revised 19.9.2011, accepted 27.9.2011.     

Taphonomic Analysis of Micromammal Remains from Striped Owl (Pseudoscops clamator) Pellets in Northeastern Buenos Aires Province,Argentina: Implications for Archaeological Sites Formation

The main objective of this study was to determine the taxonomic and taphonomic characteristics of the micromammal remains recovered from pellets of Pseudoscops clamator (striped owl), collected at three localities in northeastern Buenos Aires Province, Argentina (Punta Lara Natural Reserve, 34°49'02,6''S, 58°03'03,9''W; Costanera Sur Ecological Reserve, 34°36'6,44" S, 58°21'33,22" W; Los Robles Park, 34°40'22,03''S, 58°52'18,88''W). The main taphonomic variables (e.g. evidence of digestive action, breakage patterns and relative abundance of skeletal elements) suggest that this owl mainly produces intermediate to moderate modification. On the other hand, P. clamator preyed mainly upon large‐sized (>150 g) micromammals (e.g. Lutreolina crassicaudata, Cavia aperea, Holochilus brasiliensis and Rattus sp.), and to a lesser degree on medium‐ (50–150 g) and small‐sized species (<50 g) (e.g. Scapteromys aquaticus, Calomys sp., Oligoryzomys flavescens, Akodon azarae and Mus musculus). Fossil assemblages with a dominance of large‐sized micromammals are commonly associated with humans as agents of accumulation. However, this study demonstrates that this owl produces assemblages with abundant large‐sized micromammals, which introduces an equifinality problem. In that sense, digestive corrosion marks, breakage patterns and the relative abundance of skeletal remains are the main attributes to differentiate P. clamator from humans, as agents of accumulation. Finally, our results might serve as an analytical model for the taphonomic interpretation of the fossil record of micromammals from paleontological and archaeological sites, which fall within the distributional range and habitat of P. clamator. Copyright © 2013 John Wiley & Sons, Ltd.     

Carnivores and their prey in the Wezmeh Cave (Kermanshah,Iran): a Late Pleistocene refuge in the Zagros

Wezmeh Cave is located on the northeastern edge of the Islamabad plain, a high intermontane valley in the western‐central Zagros. In 1999 a disturbed but large faunal assemblage was recovered from this site. The abundant and extremely diverse faunal spectra present at Wezmeh Cave has highlighted the importance of this assemblage. Carnivore remains constitute the bulk of the assemblage; red fox (Vulpes vulpes) has the highest number of identified specimens followed by spotted hyena (Crocuta crocuta), brown bear (Ursus arctos), wolf (Canis lupus), felids (lion, leopard, lynx/caracal and wildcat), mustelids (badger, polecat, marten) and viverrids (mongoose). Artiodactyls (bovid, cervid, suid), equids, rhinoceros (Dicerorhinus sp.) and small animals (Cape hare, porcupine, tortoise, snake, birds) are also present. According to U‐series dating, the site was occupied from around 70 ka BP through to sub‐recent periods by carnivores. Amongst this rich assemblage, a human fossil tooth was also found and dated by non‐invasive spectrometry gamma dating to 20–25 ka BP. A preliminary zooarchaeological and taphonomic study shows that Wezmeh Cave was used by multiple carnivore species, a unique phenomenon in the Zagros Mountains in particular and southwest Asia in general. Copyright © 2008 John Wiley & Sons, Ltd.     

Lagostomus maximus (Desmarest) (Rodentia,Chinchillidae), the extant plains vizcacha in the Late Pleistocene of Uruguay

Ubilla, M. & Rinderknecht, A., April 2016. Lagostomus maximus (Desmarest) (Rodentia, Chinchillidae), the extant plains vizcacha in the Late Pleistocene of Uruguay. Alcheringa 40, xxx–xxx. ISSN 0311-5518

The extant plains vizcacha, Lagostomus maximus, is described from the Late Pleistocene (Dolores Formation) of Uruguay based on an almost complete articulated skeleton. It is compared with the nominally extinct Pleistocene species of the genus. An AMS ¹⁴C taxon-age is determined for L. maximus at 11 879 ± 95 years BP (cal. BP 13 898–13 941). Lagostomus maximus is absent from modern mammal communities in Uruguay, and no Holocene evidence is available. Because L. maximus exhibits remarkable sexual and ontogenetic dimorphism, we examined a range of juvenile, sub-adult and adult male and female specimens. It is not possible to differentiate the articulated Pleistocene fossil from sub-adult specimens of L. maximus based on this sample. Moreover, the skull characteristics, including a broad vertical ramus of the zygomatic arch, semi-circular temporal crest and short robust sagittal crest, most closely resemble extant female individuals. Molar size is non-predictive for ontogenetic stage or body mass because it reaches stability during early adulthood. Other fragmentary skull remains are herein assigned to L. sp. cf. L. maximus. Lagostomus cavifrons from the Pleistocene of Argentina is considered synonymous with L. maximus. Its character states concur with the range of intraspecific variability and it has been established on a young adult or adult male individual. Likewise, the other Argentinean Pleistocene species based on incomplete mandibles, L. angustidens, L. striatus, L. heterogenidens, L. egenus, L. minimus and L. debilis, concur with ontogenetic morphs and are here assigned to Lagostomus sp. The ecological preferences of extant L. maximus infer open arid or semi-arid landscapes for the latest Pleistocene of southern Uruguay. This hypothesis is reinforced by the coeval presence of Microcavia, Galea and small camelids in the same strata. The Last Glacial Maximum likely promoted this environmental setting. Latest Pleistocene or early Holocene climatic change might have facilitated local extinctions and/or range shifts among this mammal fauna.

Martín Ubilla [martinubilla@gmail.com], Facultad de Ciencias, Universidad de la Republica, Iguá 4225, 11400, Montevideo, Uruguay; Andrés Rinderknecht [apaleorinder@yahoo.com], Museo Nacional de Historia Natural, Paleontología, 11000, Montevideo, Uruguay.