Specialized Early Upper Palaeolithic Hunters in Southwestern France?

Paul Mellars has long used cave and rockshelter ungulate faunal assemblages from southwestern France to argue that the early Upper Palaeolithic people of this region focused their hunting on reindeer (Rangifer tarandus), and that such specialized hunting distinguishes the Upper from the Middle Palaeolithic in at least this region. We examine this argument quantitatively, using a sample of 133 Mousterian, Châtelperronian, and Aurignacian ungulate assemblages. We show that only five Aurignacian assemblages, from three sites, stand out in terms of the degree to which their ungulate faunas are dominated by a single taxon. We also show that some Mousterian cave and rock shelter ungulate assemblages are more heavily dominated by large bovids than Aurignacian assemblages are dominated by reindeer, and that Mellars' argument is highly dependent on the exclusion of open sites from the analysis and on the numerical threshold he has selected to indicate hunting specialization.     

An exclusively hyena-collected bone assemblage in the Late Pleistocene of Sicily: taphonomy and stratigraphic context of the large mammal remains from San Teodoro Cave (North-Eastern Sicily,Italy)

A detailed taphonomic analysis of the large mammal assemblage from 1998 to 2006 excavations at San Teodoro Cave is presented, taking into account the stratigraphic context of the deposits. Three not strictly contemporary fossiliferous levels having different lithological features have been detected, here named B-I, B-II, and B-III. Fossil remains are prevalently accumulated in B-I and B-II. The three levels are characterized by evidence of Crocuta crocuta spelaea occupation, represented by their skeletal remains, coprolites, and distinctive damages on the bones, similar to fossil and modern spotted hyena dens from Europe and Africa. A differential distribution of coprolites and small digested bones, probably due to different humidity conditions, has been recognized in B-I and B-II, and can be related to different topographic locations within the cave or to different climate conditions during the sedimentation phases. The very low density of fossil remains in B-III, which is the oldest level, could indicate an area that was less inhabited by hyenas, probably due to geomorphological conditions. Taphonomic comparison of the three fossiliferous levels of the San Teodoro Cave deposits points to a long-term, perhaps cyclic, occupation of the cave by hyenas and confirms the cave as one of the most important Pleistocene hyena dens in Europe.     

Chagyrskaya Cave:A Middle Paleolithic Site In The Altai

This article presents the results of multidisciplinary studies conducted at Chagyrskaya – an Upper Pleistocene karst cave in the northwestern Altai where Middle Paleolithic stone tools and fossil remains of Neanderthals were found. Taphonomic aspects of the site are addressed, and results of science-based analyses including radiocarbon and paleomagnetic dating are presented. The deposits are similar to loess-like Upper Pleistocene loams of Western Siberia. Among the Middle Paleolithic industries of the Altai, the Chagyrskaya industry is paralleled only by that of Okladnikov (formerly Sibiryachikha) Cave. Both represent a local Middle Paleolithic Mousteroid facies, named Sibiryachikha after the eponymous site.     

Camelus from the upper pleistocene of Mount Carmel,Israel

A Camelus metapodial, found among the animal bones excavated by Garrod from the Levalloiso-Mousterian of the Tabun Cave (Mount Carmel, Israel), adds to the relatively small number of Camelus finds reported from the Pleistocene of the Near East. The mammal faunas associated with these finds suggest relatively dry open conditions, but not desert conditions. The Tabun specimen is of moderate size, in contrast to the very large Camelus from Far'ah II; two species of camel may thus have been present in the Near East during the Upper Pleistocene.     

Late Middle Palaeolithic occupations in Ciemna Cave,southern Poland

Recent excavations in Ciemna Cave in the Pr?dnik valley near Ojców, southern Poland have brought to light new stratigraphic and techno-typological evidence concerning Late Middle Palaeolithic groups and their cultural affinities. In 2007, excavations began in the hitherto-unexplored main chamber of Ciemna Cave, with the goal of clarifying the results of previous work in other parts of the cave. During excavation the rocky floor of the cave was reached. About 1000 stone artifacts have been collected to date. Three cultural traditions have been documented: Mousterian, Taubachian, and Micoquian. Within the Micoquian tradition, three cultural levels were observed, which enriches the previous understanding of occupational phases at the site. These findings permit revision of the traditional terms “Pr?dnik industry” and “Pr?dnik technique.”     

The reconstruction of past environments through small mammals: from the Mousterian to the Bronze Age in El Mirón Cave (Cantabria,Spain)

El Mirón is a large cave in the Cantabrian Cordillera of northern Spain that presents a long archaeostratigraphic sequence radiocarbon-dated by over 60 assays to between 41,000 and 2000 BP. The sediments, collected from four areas within the cave and sieved-washed with fine wire meshes, contain microvertebrate remains of fish, frogs, lizards, birds and mammals, of which the latter are most abundant. Preliminary taphonomic analysis suggests that the microvertebrates were naturally collected by owls and (less) small carnivores. Small mammal assemblages are useful for palaeoenvironmental reconstruction because they are linked to particular habitats and are sensitive to environmental changes. The small mammals from El Mirón are ideal for this because sample sizes are large, bone preservation is good, and the stratigraphic sequence is long. In this paper we reconstruct the late Quaternary environments in the Cantabrian region of Spain using small-mammal assemblages from El Mirón Cave. On the basis of the ecologic adaptations of this suite of fauna, the majority still extant, we have identified seven habitat types, which are plotted through time. The evolution of the small mammal assemblages at El Mirón reveals seven major climatic shifts that correspond closely to the climatic changes recognized in the Iberian Peninsula during the last 41 kyr.     

An Early Upper Palaeolithic Stone Tool Assemblage from Mughr El-Hamamah,Jordan: An Interim Report

Mughr el-Hamamah (Jordan) Layer B contains an Early Upper Palaeolithic stone tool assemblage dating to around 39–45 kya cal b.p. This assemblage is unusual in that it samples human forager activities around the ecotone between the Transjordanian Plateau and the palaeo-lake (Lake Lisan) that filled much of the Jordan Valley during Late Pleistocene times. This paper describes that assemblage, comparing it to other Levantine Upper Palaeolithic assemblages of equivalent antiquity. The Mughr el-Hamamah Layer B assemblage appears most similar to Early Ahmarian assemblages, but it departs from typical such assemblages in ways that may reflect local conditions’ influence on human activities carried out in and near the cave. Mughr el-Hamamah raises new questions about changes in residential mobility, off-site provisioning and foraging activity, and on-site task diversity in the Early Upper Palaeolithic period.     

On the occurrence of the scimitar-toothed cat, Homotherium latidens (Carnivora; Felidae), at Kents Cavern,England

Teeth of Homotherium latidens recovered from late Pleistocene sediments, Kents Cavern, England have long been the source of controversy. H. latidens is conspicuously absent from other late Pleistocene cave deposits in Britain, and is widely thought to have been extirpated from the region during the isotope stage 10 glacial period. Here we present high spatial resolution analyses of fluorine and uranium uptake profiles in teeth of three species from the same cave. The H. latidens tooth is clearly distinguished from the unambiguously provenanced Late Pleistocene hyaena and Middle Pleistocene cave bear teeth. These results are consistent with the theory that the H. latidens teeth originated at an exogenous location, were probably transported to Kents Cavern as Palaeolithic trade goods, and were buried in Kents Cavern in Palaeolithic times.     

An Early Pleistocene Phytolith Record from Wonderwerk Cave,Northern Cape,South Africa

Grass silica short cell phytoliths were sampled from the four lowermost archaeological strata in excavation 1 at Wonderwerk Cave and offer an independent record of climatic change during an episode of Early Stone Age hominin occupation at the cave. Linked to differences in growing season temperature and the geographic distribution of C₃ and C₄ grasses in southern Africa, fossil grass phytoliths were used to trace palaeoenvironmental shifts at the site. The results suggest that Early Pleistocene environmental conditions at the cave fluctuated: between wetter and drier summer–rainfall growing conditions (C₄) towards the end of the Olduvai subchron and the beginning of the subsequent interval of reversed polarity, to mostly dry and cooler winter–rainfall growing conditions (C₃), that continued throughout the interval. It ended with a shift towards increased summer rainfall aridity at around one million years ago. The fluctuation between markedly wetter and drier C₄ conditions at the cave (NADP-me grass types vs. NAD-me grass types) does not support the premise that the expansion of C₄ grasslands was always coupled with increased aridity.     

Implications of age variation and sexual dimorphism in modern equids for Middle Pleistocene equid taxonomy

Interspecific variation in the equid lineage is limited when compared with other large mammal lineages, while a considerable amount of intraspecific ecomorphological variation occurs. This has led to considerable debate regarding the taxonomic status of palaeontological and archaeological equid remains. The possible confounding impact of sex and age variation on analyses of archaeological equid remains has so far remained untested. This could be especially problematic when it is unknown whether an assemblage contains more than one equid species, breed or hybrid. Here, sex and age variation in modern equid species is examined and compared to variation in Pleistocene equid assemblages from northwest Europe. Sex variation in the longbones of modern equid species was found to be lower than variation over time. Similarly, age variation in dental elements of the same species is predominantly lower than diachronic variation, although caution needs to be taken when an archaeological assemblages consists mostly of elements at a similar wear stage. To assess whether late Middle Pleistocene caballoid horse fossils should be assigned to multiple species or be considered to form a monospecific group, the coefficients of variation for measurements on fossil horse remains were compared with those for modern equid specimens. Results indicate that the fossil assemblages are similarly variable to samples of modern material. Taken as a whole, the late Middle Pleistocene caballoid horse lineage is equally variable to an assemblage of modern ponies of a variety of different breeds. This implies that the variation observed between samples dating from different parts of the Middle Pleistocene can be characterised as intraspecific variation within a single species reflecting ecomorphological adaptations to the changeable Pleistocene climate. Copyright © 2010 John Wiley & Sons, Ltd.     

U-series Dating of Fossil Teeth and Carbonates from Snake Cave,Thailand

Snake Cave (Thailand) is an important site for the study of the evolution in Southeast Asia, leading to a discovery of rich and various faunas, including a human tooth. Fossils of large and small mammals converge to indicate, on the basis of faunal assemblage, a late Middle Pleistocene age for the fossiliferous layers; on the other hand, previous U-series dating of carbonates furnished a lower limit for the fossils of about 130 ky.In order to achieve a better comprehension of the chronology of the cave, the Uranium-series method for absolute age determination has been applied to some carbonates older than those studied previously and to fossils themselves.We used the isochron technique approach to yield valuable ages because spleleothem samples consist in a mixture of calcite and detrital contamination. The results obtained suggest that the main fossiliferous level is older than about 160 ky. The application of the U-series method to tooth samples from this tropical cave leads to erroneous ages, probably because of post depositional groundwater movements.     

The Exploitation of Plant Resources by Neanderthals in Amud Cave (Israel): The Evidence from Phytolith Studies

The depositional environments of Amud Cave indicate that phytolith assemblages retrieved from the cave's sediments are an integral part of the Middle Palaeolithic sequence. As such, they provide direct evidence for plant use. The Amud Neanderthals emphasized both wood and grass exploitation. Ligneous parts of trees and shrubs were used mainly for fuel. Herbaceous plants were used for bedding, possibly fuel, and for food. There is clear and repetitive evidence for the exploitation of mature grass panicles, inferred to have been collected for their seeds. These findings suggest that, as with the pattern recently discerned for faunal resources, a broad spectrum of plants has been exploited from at least the end of the Middle Palaeolithic. Phytolith analysis now provides a tool for testing models explaining subsistence and mobility patterns during the Levantine Middle Palaeolithic and for better understanding the role of vegetal resources in shaping these patterns.     

Taphonomy and zooarchaeology of the Upper Palaeolithic cave of Dzudzuana,Republic of Georgia

We present the results of a detailed taphonomic and zooarchaeological study of the faunal remains from the Upper Palaeolithic layers of Dzudzuana Cave, Republic of Georgia. This study presents the first carefully analysed Upper Palaeolithic faunal assemblage from the southern Caucasus and thus serves as a significant point of reference for inter‐regional studies of Upper Palaeolithic subsistence in Eurasia. A series of intra‐site taphonomic comparisons are employed to reconstruct the depositional history of the bone assemblages within the different occupational phases at the site and to investigate subsistence, meat procurement and bone‐processing strategies. Caucasian tur (Capra caucasica), aurochs (Bos primigenius) and steppe bison (Bison priscus) were the major prey species throughout the Upper Palaeolithic. Their frequencies do not change significantly over time, and nor does bone preservation vary by layer. The assemblage is characterised by significant density‐mediated biases, caused by both human bone‐processing behaviours and in situ post‐burial bone attrition. Bone marrow extraction produced large numbers of unidentified bone fragments, many exhibiting green bone fractures. The density and size of bone assemblages and the extent of fragmentation indicate that Dzudzuana Cave was repeatedly occupied by Upper Palaeolithic foragers over many years. Skeletal part representation and butchery marks from all stages of carcass processing suggest that prey occasionally underwent field butchery. Intra‐site taphonomic comparisons highlight uniform patterns of cultural and economic behaviours related to food procurement and processing strategies. Copyright © 2007 John Wiley & Sons, Ltd.     

Human consumption of tortoises at Level IV of Bolomor Cave (Valencia, Spain)

The acquisition and consumption of small prey in the pre-Upper Palaeolithic is a highly debated topic at present. For some authors, the systematic obtaining of these animals is only part of the subsistence strategies used by anatomically modern Humans. However, the consumption of small prey dates back to the Plio-Pleistocene chronologies in some sites. Although the utilization of leporids has been recorded in several pre-Late Pleistocene European sites, the evidence of tortoise consumption is documented not as common for these periods. However, Level IV of Bolomor Cave has clear diagnostic elements to document the acquisition and use of tortoises (Testudo hermanni) for food in the form of: (1) cutmarks on limb bones and ventral surface of the carapace and plastron; (2) presence of burning on tortoise skeleton and shell; (3) elements of anthropogenic breakage on carapace and plastron: percussion pits, percussion notches and impact flakes; and (4) human toothmarks on limb bones. This paper tries to examine the possible patterns in the tortoise consumption sequence from Level IV of Bolomor Cave and improves data on the butchery process and tortoise consumption in the Late Middle Pleistocene.     

Middle Pleistocene bird consumption at Level XI of Bolomor Cave (Valencia, Spain)

The consumption of small prey dates back to the Plio-Pleistocene chronologies in some African sites. However, the systematic acquisition and consumption of small prey in the pre-Upper Palaeolithic times is still a highly debated topic in Europe. Although the utilization of leporids has been recorded in several pre-Late Pleistocene European sites, the evidence of bird consumption is not as common for these periods. Nevertheless, Level XI (MIS 6) of Bolomor Cave has clear diagnostic elements to document the acquisition and use of birds (Aythya sp.) for food in the form of: (1) cutmarks on bones of both the front and hind limb; (2) presence of burning patterns on the extremities of the bones (areas of the skeleton with less meat); and (3) human toothmarks on limb bones. The capture of birds is classified as quick-flying game in the archaeological sites. The acquiring of fast-running (mostly lagomorphs) and quick-flying small prey requires a sophisticated technology and involves obtaining and processing ways different from those used for large- and medium-sized animals. From this perspective, the aim of this paper is to examine possible patterns in the processing sequence of birds from Level XI of Bolomor Cave and to improve the data on their butchery and human consumption in the Middle Pleistocene of Iberian Peninsula.     

Probable evidences of bone accumulation by Pleistocene bearded vulture at the archaeological site of El Mirón Cave (Spain)

The bearded vulture (Gypaetus barbatus), a bird or prey that mainly feeds on bones of large mammal carcasses, has been recently evidenced as an accumulating agent with significant contributions to palaeontological assemblages in caves and shelters, but extremely rare at archaeological sites. The results of the taphonomic analyses carried out in the Upper Palaeolithic layers of El Mirón Cave (Cantabrian Spain), which are presented here, evidence the existence of special digestive marks and a typical skeletal pattern in a noticeable amount of small and medium-sized ungulates bones. This fact suggest that a representative part of the bone assemblage has been caused by bearded vultures, and strengthens the idea that this bird has to be definitely included among other potential non-human accumulators in archaeological sites.     

Identifying palaeo-environments and changes in Aboriginal subsistence from dual-patterned faunal assemblages,south-western Australia

Palaeo-environments and past human subsistence patterns are difficult to determine from dual-patterned faunal assemblages where human and non-human predators have accumulated and intensively modified animal bones. This paper examines such records in the Leeuwin–Naturaliste Region of south-western Australia, where a thin belt of coastal limestone contains caves and rock shelters with rich faunal deposits. The Late Pleistocene and Holocene part of this record derives from four archaeological sites: Devil's Lair, Tunnel Cave, Witchcliffe Rock Shelter and Rainbow Cave. Correspondence analysis combined with cluster analysis enables a preliminary assessment of habitat changes using simple species abundances in the faunal assemblages and comparison with indices of past human activity in the sites and the species’ present habitat preferences. These inferred changes, consistent with previous analyses of faunal remains and tree charcoal, suggest that late Holocene sites document Aboriginal occupation in coastal heath, scrub and woodland. Late Pleistocene deposits record hinterland occupation at times of low sea-level when the coast was up to 30 km seawards of its present position and the surrounding vegetation was open-forest or woodland. As rainfall increased and vegetation changed in the Holocene, species foraging in open-woodland declined or became locally extinct, while species requiring closed canopy habitats increased. Rank-order correlations of taxa and archaeological remains from depositional sequences before and after the environmental change indicate that the occupiers of late Holocene sites favoured the same generalist species that occupiers of Late Pleistocene sites had favoured, which were available at all times. Prey habitats, foraging behaviours and historic records of ethnographic hunting and settlement pattern suggest that this local continuity is consistent with maintenance of a “dispersive mode” subsistence pattern in the region.     

Mammalian fauna from Palaeolithic sites in the Upper Yenisei River basin (southern Siberia): Review of the current zooarchaeological evidence

The palaeontological data on mammal remains from two Middle Palaeolithic (ca. 125 000 years ago – 27 200 BP) and 34 (ca. 34 000–12 000 BP) Upper Palaeolithic cultural complexes in the upper part of the Yenisei River basin (southern Siberia) are considered. General features of the faunal assemblages are established. Several issues are discussed, including (a) changes in species composition through time and palaeoenvironmental implications of the zooarchaeological records; (b) patterns of human exploitation (hunting) of mammals; (c) issue of mammoth hunting and (d) possibility of domestication of dog in the late Upper Palaeolithic in the Yenisei River basin. Copyright © 2009 John Wiley & Sons, Ltd.     

The British Late Middle Palaeolithic: An Interpretative Synthesis of Neanderthal Occupation at the Northwestern Edge of the Pleistocene World

The British Middle Palaeolithic is divided into two discrete periods of occupation: the Early Middle Palaeolithic (MIS 9–7, ~330–180 ka BP) and the Late Middle Palaeolithic (MIS 3, ~59–36 ka BP), separated by a long hiatus. Owing to the relative poverty of the record and historical difficulties in dating and correlating archaeological sites, the British Late Middle Palaeolithic has, until recently, received scant attention, and has largely been regarded as the poor man of Europe, especially by British archaeologists. Indeed, there has been more discussion of the absence of humans from Britain than of what they did when they were present. We aim here to redress that situation. Following from recent considerations of the Early Middle Palaeolithic (White et al. in J. Quat. Sci. 21:525–542, 2006; Scott, Becoming Neanderthal, Oxbow, Oxford, 2010), we offer an interpretative synthesis of the British Late Middle Palaeolithic, situating ‘British’ Neanderthals in their chronological, environmental and landscape contexts. We discuss the character of the British record, and offer an account of Neanderthal behaviour, settlement systems and technological practices at the northwestern edge of their known Upper Pleistocene range. We also examine the relationship of the enigmatic Early Upper Palaeolithic leafpoint assemblages to Neanderthals.     

Towards a Chronology of the Jerzmanowician—a New Series of Radiocarbon Dates from Nietoperzowa Cave (Poland)

The period around 50 000–35 000 years ago constitutes one of the most debated research issues in European archaeology of the Palaeolithic. In this time period, the transition from the Middle to the Upper Palaeolithic took place. Locally, in different areas of Europe, this shift is recorded in so‐called transitional assemblages. The eastern fringe of these transitional assemblages is represented by the Jerzmanowician, the unit described on the basis of a lithic assemblage from Nietoperzowa Cave (Poland). The unit is a part of a European transitional complex called the Lincombian–Ranisian–Jerzmanowician. Up to now, the radiocarbon dates presented in the literature have only allowed us to set the age of the Jerzmanowician at c.40 000–45 000 ka cal bp . In this study, we present 42 new radiocarbon dates. We attempt to set the archaeological record of the Jerzmanowician from Nietoperzowa Cave in an accurate chronological framework, based on Bayesian statistical processing of radiocarbon dates. We conclude that the lower boundary of layer 6 in Nietoperzowa Cave can be statistically located in the range 44 000–42 000 cal bp and the upper limit for the Jerzmanowician is estimated to c.31 000 cal bp . New data raises a question on the correlation with upper layer 4. In the light of the new chronology, the attribution of the archaeological inventory from layer 4 to the Jerzmanowician seems questionable.