Hiding to eat: the role of carnivores in the early Middle Pleistocene from the TD8 level of Gran Dolina (Sierra de Atapuerca,Burgos, Spain)

Damage generated by large and small carnivores is common in many Middle Pleistocene sites. However, identifying the predator that produces the faunal accumulations is often a difficult task. In order to recognize the main type of carnivore that acts on a faunal assemblage, a combination of several characteristics should be taken into account: taxonomic and skeletal element representation, age profiles, carnivore damage (location, frequencies and dimensions of tooth-marks, bone breakage and digested bones), degree of fragmentation and frequencies of coprolites. But, adding environmental characteristics and the ethology of non-human predators/scavengers is also important. All these aspects are applied to the faunal assemblage from the TD8 level of the Gran Dolina site (Sierra de Atapuerca, Burgos, Spain). Paleomagnetic data combined with ESR and U-series place the TD8 level at the beginning of the Middle Pleistocene, specifically circa 700 kyr ago. The TD8 level contains a large faunal accumulation primarily composed of ungulate skeletal elements, and to a lesser extent carnivore remains. This assemblage is characterized by an overrepresentation of fallow deer (Dama vallonetensis), a skeletal profile biased towards cranial remains and limb bones, diversity of ages at death, a high proportion of carnivore damage and tooth-marks of large size, and an absence of human activity. According to these data, the accumulation seems to have been produced primarily by large carnivores, possibly hyenas. This observation does not rule out the possible occasional activity by other carnivores. Nevertheless, the characteristics of the TD8 assemblage do not correlate entirely with those traditionally used to define carnivore dens. In TD8, there are (1) no immature carnivore remains (remains of just one young Mosbach wolf); (2) scarce traces related to the end stages of consumption and some anatomical connexions; (3) few coprolites; (4) high proportion of adult ungulates and; (5) high quantities of whole bones and epiphyses. From this perspective, the TD8 faunal assemblage seems to correspond to a succession of carnivore occupations that allows the development of a suite of features to identify the activities of several species of predators that may have used the cave in different ways and durations. This study aims to emphasize the importance of these analyzes in order to know the behaviour of different non-human predators/scavengers in the European Middle Pleistocene sites.     

Spatial organization of the Gravettian mammoth hunters' site at Kraków Spadzista (southern Poland)

One of the largest and best known sites of the Central European Gravettian is the Kraków Spadzista site. This article summarizes archaeological, palaeontological, and zooarchaeological data from the site, obtained in the course of more than 40 years of excavations and studies. The site is known mainly from the discovery of the large mammoth bone accumulation in trench B. During field work conducted in a few trenches located close to this assemblage, numerous human artifacts and faunal remains were found. On the basis of analyses of lithic materials, faunal remains, and radiometric data it is concluded that the Krakow Spadzista site was frequently re-occupied during a few weeks or months, by groups of Gravettian hunters in the period 24–22,000 rcyBP This short-term camp site was related to mammoth hunting. The analysis allowed identification of three possible activity zones, characterized by the presence of different groups of flint tools, fireplaces, and different faunal assemblages: a camp area, a lithic workshop, and an animal-processing area, besides the well known accumulation of mammoth bones.     

Museum Collections and Archaic Era Vertebrate Faunal Remains from Cuba

ABSTRACT

Museum collections are an underutilised source of environmental archaeological data. Curated collections often contain faunal and floral remains that can be analysed using new methods and to address various research questions. For example, the broad patterns of Archaic or preceramic subsistence adaptations are not well known for the Caribbean region, and for Cuba in particular, due to the ecological variability among the islands and limited, quantified faunal data from radiocarbon-dated sites. To address these issues, we present quantified vertebrate faunal data and new radiocarbon dates from museum-curated collections from three Cuban sites. Las Obas, La Vega del Palmar, and Los Caracoles are Archaic, dating between roughly 350 BC to AD 630, although Vega del Palmar contains Pre-Arawak Pottery Horizon ceramics. The three assemblages include a diversity of taxa from several habitats, including various species of hutia (Capromyidae). The taxonomy of these rodents varies among the sites as do the proportions of other taxa. In addition to abundant hutia, the sites' occupants exploited an array of fish, birds, reptiles and other mammals, including manatee. We compare the sites faunal assemblages and discuss their ecological significance as well as their place in Cuban prehistory.     

Variability in Faunal Assemblages: The Influence of Hunting Skill, Sharing, Dogs, and Mode of Cooking on Faunal Remains at a Sedentary Kalahari Community

Variability in faunal assemblages from different sites and/or from different time periods is often attributed to economic or taphonomic factors. The role of sharing on faunal remain distributions is compared to other factors that have been suggested to influence these distributions, such as hunting skill. Faunal species and skeletal elements are compared among three hunter—gatherer camps that form a sharing network. These are contrasted with those of two other hunter—gatherer camps located at the same Kalahari community occupied by an unusual family that is a relative isolate in terms of sharing. The effect of sharing on equalizing variation in hunting success as reflected in the faunal remain inventory is explored from the five camps inventoried in 1990. Complicating factors which tend to affect faunal remain frequencies are also examined, such as cooking technique and dogs. All faunal remains visible on the surface of each camp were recorded according to species, element or fragment portion, age (mature or immature), and, when possible, side. At all but one camp, surface faunal remains were recorded both before and after ethnographic observations during the dry season of 1990. In addition to hunting success, all occurrences of sharing and consumption of meat were recorded during these observation periods and those conducted on and off between 1987 and 1992. Although participation in a sharing network obscures differences in hunting skill in the archaeological record, sharing impacts on faunal assemblages in interesting ways that are potentially archaeologically visible. Sharing in strongly egalitarian societies levels unequal hunting skill that could otherwise affect faunal remain frequencies, taxa richness, meat weight, and other indices measured here. In these societies, sharing reinforces social bonding between kin and friends in ways that help unit families from different camps.Zooarchaeologists have become accustomed to high levels of confidence in their inferences about the origins, functions, and responses to stress of animal remains. This confidence rests on the causal and functional links between attributes of these remains and the processes and contexts which generate them. Their investigations are presently moving toward wider inferences about the context and functions of bones in ancient hominids' behavioral systems… Zooarchaeologists now need a different set of inferential strategies than that which characterized their preceding phase of research. (Gifford 1991:215)     

Neandertal foraging during the late Mousterian in the Pyrenees: new insights based on faunal remains from Gatzarria Cave

This article presents the initial results of a new study of faunal remains from Gatzarria Cave, a Middle-to-Upper Palaeolithic transition site the Pyrenees of southwestern France (department of the Pyrénées Atlantiques). This study attempts to document diet breadth during the late Mousterian, while paying due attention to recently identified problems regarding the stratigraphic context of the assemblages. The faunal analysis focuses on a subset of late Mousterian faunal remains from layer Cj at the site. Taphonomic analysis suggests that humans were the primary bone accumulators. The assemblage is dominated by a single large-bodied species, red deer; smaller-bodied ungulates are poorly represented. Skeletal part representation indicates that within-bone nutrients contained in marrow were probably a key resource for these foragers. The overall pattern of remains is interpreted as evidence of narrow-spectrum foraging, a pattern which appears to be repeated at other Mousterian sites in the Pyrenees region. This may mean that local Neandertal populations existed at relatively low densities. However, this suggestion must be tempered by the fact that settlement patterns, including occupation seasonalities and site functions, are not yet well understood for this region.     

Middle Holocene fishing strategies in East Africa: zooarchaeological analysis of Pundo,a Kansyore shell midden in northern Nyanza (Kenya)

A zooarchaeological and taphonomic analysis is presented for the faunal remains from the Pundo shell midden in northern Nyanza province (Kenya), which contains Kansyore ceramics and dates to ca. 8000–7600 cal. BP. The faunal assemblage is overwhelmingly dominated by fish and molluscs, of which only the fish data are presented in detail. Taxonomic identifications suggest a diachronic change in relative species abundance, with cichlids becoming increasingly dominant, eventually comprising nearly 80% of the fish assemblage. The Pundo faunal data are compared with those from a number of other Kansyore sites. Pundo confirms that early Kansyore sites are purely forager sites, with domestic caprines only appearing at late Kansyore sites (post‐dating ca. 4400 cal. BP). Comparing ethological data for fish taxa found at Pundo and other lakeshore shell middens with those for fish taxa found at larger riverside sites, a seasonal round is suggested, in which short‐term fishing camps such as Pundo may have been occupied in the dry season, while riverside sites were probably occupied during the rainy season. Copyright © 2008 John Wiley & Sons, Ltd.     

Isotopic palaeodiet studies of Ancient Egyptian fauna and humans

We report here stable carbon and nitrogen isotope results from human and faunal bone collagen from samples taken from sites in the Egyptian Nile Valley and surrounding region, dating from the Predynastic (c. 5500 BC) through to the Dynastic (c. 343 BC) periods. Isotopic values for the human population cluster together across this broad time range, with high δ¹⁵N ratios, and δ¹³C values indicating a largely C₃ based diet. The human data is not easily explained through comparison with our associated faunal data, and so may be explained by the consumption of protein from an ecosystem we did not adequately sample, such as freshwater fish or plants and fauna with unusually high δ¹⁵N values due to the extreme aridity of this region. The faunal isotopic data we did produce shows a great range in values between and within species, especially in cattle, and reflects the close proximity of three ecozones across a relatively narrow geographical area; the river, the immediate flood-plain area around the Nile and the desert surroundings.     

Identifying palaeo-environments and changes in Aboriginal subsistence from dual-patterned faunal assemblages,south-western Australia

Palaeo-environments and past human subsistence patterns are difficult to determine from dual-patterned faunal assemblages where human and non-human predators have accumulated and intensively modified animal bones. This paper examines such records in the Leeuwin–Naturaliste Region of south-western Australia, where a thin belt of coastal limestone contains caves and rock shelters with rich faunal deposits. The Late Pleistocene and Holocene part of this record derives from four archaeological sites: Devil's Lair, Tunnel Cave, Witchcliffe Rock Shelter and Rainbow Cave. Correspondence analysis combined with cluster analysis enables a preliminary assessment of habitat changes using simple species abundances in the faunal assemblages and comparison with indices of past human activity in the sites and the species’ present habitat preferences. These inferred changes, consistent with previous analyses of faunal remains and tree charcoal, suggest that late Holocene sites document Aboriginal occupation in coastal heath, scrub and woodland. Late Pleistocene deposits record hinterland occupation at times of low sea-level when the coast was up to 30 km seawards of its present position and the surrounding vegetation was open-forest or woodland. As rainfall increased and vegetation changed in the Holocene, species foraging in open-woodland declined or became locally extinct, while species requiring closed canopy habitats increased. Rank-order correlations of taxa and archaeological remains from depositional sequences before and after the environmental change indicate that the occupiers of late Holocene sites favoured the same generalist species that occupiers of Late Pleistocene sites had favoured, which were available at all times. Prey habitats, foraging behaviours and historic records of ethnographic hunting and settlement pattern suggest that this local continuity is consistent with maintenance of a “dispersive mode” subsistence pattern in the region.     

Animal Resource Exploitation among the Virgin River Puebloans in the American Southwest

Abstract

A review of the literature documenting faunal remains recovered from 16 archaeological sites attributed to the Virjin River Puebloan (VRP) tradition (ca. 300 B.C.–A.D. 1200) of the American Southwest was conducted to identify the importance of wild animal resources to the subsistence strategy. The Number of Individual Specimens (NISP) was counted (NISP = 16,386) and sorted by taxa and animal size categories to explore differences in the distributions of faunal remains by archaeological recovery strategy, site type, physiographic location, and chronology. The most significant factor in the differential use of animal resources among the VRPs was physiographic location. Desert lowland sites had greater species diversity. Sites in the intermediate St. George Basin had more large mammals such as deer, and sites located on the higher plateau had more small mammals such as lagomorphs. These results indicate that the subsistence strategy varied across diverse environmental conditions, but animal resources were always an integral part of the VRP adaptation until their abandonment of the region in the early 13th century A.D.     

The mobile herding economy of Kebarah Cave,Mt carmel: An economic analysis of the faunal remains

The faunal remains from Kebarah Cave provide evidence of the economic activities of the cave's inhabitants during the late Pleistocene and early Holocene. The variety of grazing resources around the cave is reflected in the proportions of the several species exploited, which changes gradually over time. A comparison of Bos remains indicates that locational factors appear to outweigh any chronological tendency towards size reduction. It is suggested that the same may be the case for animals not usually associated with human exploitation, in particular the hyaena. The data from an excavation conducted 40 years ago are inadequate to answer the questions posed by bioarchaeologists today, but the evidence from better known neighbouring sites has subsequently borne out much of what could be proposed here as hypothesis.     

Animal engravings in the central Sahara: A proxy of a proxy

The rock art of the central Sahara was created out of the beliefs, traditions and experiences of the engravers and painters. The animal engravings of the Wadi al-Ajal in south-western Libya are used to isolate some of the environmental and cultural/symbolic components that make up the pictorial record. A comparison between the depicted animals and the faunal remains recovered in the area identifies a number of characteristics. The engraving repertoire is dominated by a small number of frequently depicted animals that were of symbolic or economic importance to the engravers. Rare and singular depicted species extend this record to a diverse species spectrum. Although a preferential depiction of herbivores is evident in the data, there is a close match between faunal record and engravings which shows that all larger animals (over ca. 10 kg) that were present in the area were also depicted. The selection of animals in the rock art appears to reflect their visibility in the landscape while also providing a record of changing climatic conditions from savanna to desert habitats. The rock art therefore provides an indirect record of the local environment while also capturing the engravers' perception of animals and landscape.     

陕西横山县大古界遗址动物遗存分析

大古界遗址位于陕西榆林市横山县雷龙湾乡沙峁村张油坊组,2010年发掘时出土了大量的动物遗存。按照出土单位对所有的动物骨骼进行了科学的收集以及分类、测量和鉴定。通过系统的分类和研究表明至少代表12个属种,包括蚌类、喜鹊、鹤、草兔、狗、貉、狗獾、黄鼬、猪、鹿科、狍、绵羊等。根据对出土动物骨骼的分析结果表明:当时遗址周围的环境以草原为主,不远处有一定面积的疏林、灌丛,草原、疏林间分布着一定面积的水域。家养动物猪和狗占的比例较少,主要以捕猎野生动物野兔和狍子为肉食来源。     

Ohalo II man—unusual findings in the anterior rib cage and shoulder girdle of a 19000-year-old specimen

The analysis of the skeletal remains of Ohalo II man, aged 30–40 years at death and dated to 19 000 years BP, shows advanced and highly unusual ossification of the lower costosternal cartilage; the right humerus is morphologically larger and considerably more robust than the left; degenerative changes were noted unilaterally in the right glenohumeral, acromioclavicular and claviculosternal joints. There is a marked asymmetry of the atlas, axis and occipital condyles. The changes in the costochondral area of the lower anterior rib cage are considered to represent an infectious chronic osteomyelitic process. The marked discrepancy in size between the left and right shoulder girdles and humeri, and the evidence of degenerative disease exclusively on one side only may be the result of a traumatic brachial plexus nerve palsy. The asymmetrical atlas and axis are most probably anatomical variants that do not reflect clinical pathology.     

BUTCHERING PATTERNS AND SPATIAL DISTRIBUTION OF FAUNAL ANIMAL REMAINS CONSUMED AT THE LOS MILLARES CHALCOLITHIC SETTLEMENT (SANTA FE DE MONDÚJAR,ALMERÍA,SPAIN)

The archaeological settlement of Los Millares (Santa Fe de Mondújar, Almería) is located in the Mediterranean coastal area of southern Spain. This archaeological site constitutes one of the most important settlements belonging to the Copper Age in the Iberian Peninsula and has given its name to the ‘millariense’ culture. The settlement has been completely excavated, yielding major archaeological faunal remains, with 27,426 bone fragments classified by researchers from Munich University (Germany). In this paper, we apply different statistical methods (mainly χ² analysis) to study the spatial distribution of bones from domestic and wild animals, reconstructing the patterns of appearance within specific functional spaces. These functions can be associated with meat consumption (sites to cut the animals into small pieces, areas where the meat was eaten, garbage sites, etc.) and the production of handmade objects (bone tools, stone tools, etc.). The results show different spatial distributions of faunal remains within the areas of the settlement, according to the animal species and particular parts of the animal skeleton, as well as the age and the sex of individuals. The findings reflect social behaviour that relates the consumption of certain species to a specific sector of population, and such behaviour can be discerned in the animal remains. Moreover, there is evidence regarding economic matters, such as meat production and the exchange of animals, which require further research.     

陕西高陵杨官寨环壕西门址动物遗存分析

一、前言杨官寨遗址位于陕西高陵县姬家乡杨官寨村四组东侧泾河左岸的一级阶地上,南距泾河约1公里,海拔498.5米。遗址北起雷村、南至韩村、西自杨官寨村西、东达徐吾村,南北宽约800米、东西长约1000米,面积约80万平方米。2008～2010年,陕     

Understanding faunal contexts of a complex Tell: Tel Dor, Israel, as a case study

The complex stratigraphy of the large Levantine tells and the complexity of human behavior that took place on them, poses a major challenge in understanding site formation processes and their reflection in the faunal remains. We studied the contextual deposition of faunal remains in Tel Dor, as a model for complex tell sites, and the possibility of using faunal remains as a tool to distinguish between context types. In addition, we asked how can we use this knowledge to elucidate site formation processes. Our results demonstrate that most loci defined in the field as primary refuse or purposive disposal are indeed different from the loci defined as secondary refuse. Different types of contexts can be differentiated, to a degree, from one another based upon multivariate analysis of faunal remains. Statistical as well as spatial analyses may help elucidate site formation processes and the use of space. Bones can, and in many cases do, reflect primary activities. Lumping zooarchaeological data into a single ‘assemblage’, as done in most zooarchaeological studies today causes major loss of information. Consideration of the specific location of faunal remains can be used as further indication for context identity and for understanding specific activities in a site, with care this can be done even in complex sites such as the ‘urban mounds’ of the Levant.     

陕北靖边五庄果墚动物遗存及古环境分析

五庄果墚遗址位于陕西省靖边县黄蒿界乡小界村西北,距离靖边县城约30公里.遗址面积约30万平方米以上,现为省级文物保护单位.     

Stable isotope analysis of prehistoric populations from the cemeteries of the Middle and Lower Dnieper Basin,Ukraine

This paper presents the results of new stable isotopes (carbon and nitrogen) analysis of human, faunal and fish remains from thirteen cemeteries from the Middle and Lower Dnieper Basin, Ukraine. The results are integrated with earlier analyses, undertaken solely on human samples, in order to provide a comprehensive overview of subsistence across the Upper Palaeolithic through to Eneolithic periods in this region. This is the first time that a combined sample of human, faunal and fish remains has been studied in order to interpret subsistence strategies across these periods in Ukraine. The total dataset comprises 113 samples of human, faunal and fish remains, 59 of which include new analyses of faunal and fish remains that have not previously been made available for study (Table 1). The analysis of the faunal and fish remains allows for a consideration of trophic levels which indicates that the consumption of freshwater fish occurs from the Epipalaeolithic period onwards. Whilst the majority of the cemeteries and the individuals therein cluster in relatively tight groups, there is a significant offset between the human and faunal samples due to the consumption of freshwater resources. The fish samples analysed in the study appear to exhibit a relatively random distribution when compared to the other samples analysed. In general, freshwater resources remain significant through until the end of the Neolithic period and into the Eneolithic period at the sites investigated.     

STABLE ISOTOPE ANALYSIS AND DIET IN EASTERN OKLAHOMA

Abstract

Over the last three decades, significant quantitative information on prehistoric diets has come to light from a variety of locations in the Caddoan archaeological area, a region encompassing eastern Oklahoma, western Arkansas, northeast Texas, and northwest Louisiana. Most of this research is based on macro-botanical and faunal remains analysis. As an additional line of evidence for the growing body of botanical and faunal data, stable carbon and nitrogen isotopes in bone collagen were analyzed from 82 human and faunal samples from the Arkansas basin and Ozark Highlands in eastern Oklahoma. The research investigates the role of maize in diets through time and across sex and status dimensions. Spanning six phases, within the time range 300 B.C. to A.D. 1650, mean δ¹³C values increase from –19.2 to –12.7 parts per million ‰. Although present prior to A.D. 1000 substantial increases in the use of maize do not begin until the Harlan phase (A.D. 1050–1250). Even with increased use, maize never dominates the diet to the extent seen in Mississippian period sites farther east.     

“Measure for measure”: a taphonomic reconsideration of the Kebaran site of Ein Gev I,Israel

We present the results of a detailed taphonomic study of the faunal remains from Ein Gev I, a Kebaran open-air site near the Sea of Galilee, Israel. In an earlier study faunal remains were assigned to bone element and species using a rapid method technique based on the identification of teeth and epiphyses only (Davis, 1974). In this study we identified the maximum number of skeletal elements, including fragments of long bone shafts. We also employed a series of taphonomic analyses to reconstruct the depositional history of the bone assemblage and to investigate subsistence, meat procurement and bone processing strategies. We compared the results obtained by a complete bone sampling method with those derived using a rapid method. The bone assemblage is dominated by mountain gazelle (Gazella gazelle), Mesopotamian fallow deer (Dama mesopotamica) and bezoar goat (Capra aegagrus). The assemblage is characterized by significant density-mediated biases, caused by both human bone processing behaviours and in situ post-burial bone attrition. Our attempts to compare species abundance derived from the two different identification protocols revealed no significant difference between the two sampling methods, both in terms of species diversity (richness and evenness) and skeletal element abundance (SEA). These comparisons contrast previous studies that expected to find ubiquitous head-and-foot dominated SEA profiles in zooarchaeological studies that were based on the incomplete identification of faunal remains.