Small prey hunting technology and zooarchaeological measures of taxonomic diversity and abundance: Ethnoarchaeological evidence from Central African forest foragers

Archaeological analyses of faunal assemblages often rely on rationale derived from the prey choice model to explain temporal and spatial changes in taxonomic measures of diversity and/or abundances. In this paper, we present analyses of ethnoarchaeological observations and bone assemblages created by Central African Bofi and Aka forest foragers which show that different small prey hunting technologies target specific suites of prey and that hunters vary their technological choice depending on their foraging goals. Analysis of ethnoarchaeological bone assemblages produced by the Bofi and Aka shows that variability in target prey can create spatially distinct, but contemporaneous, faunal assemblages with different diversity values and abundance indices. These data reveal important variation in how individuals within a contemporary human population rank prey and challenge current assumptions about the meaning of diversity and abundances measures in archaeological contexts. We argue that the use of diversity and abundance indices can obscure important intrasite variability in prehistoric foraging effort and suggest strategies that might enhance current techniques.     

An Early Upper Palaeolithic Stone Tool Assemblage from Mughr El-Hamamah,Jordan: An Interim Report

Mughr el-Hamamah (Jordan) Layer B contains an Early Upper Palaeolithic stone tool assemblage dating to around 39–45 kya cal b.p. This assemblage is unusual in that it samples human forager activities around the ecotone between the Transjordanian Plateau and the palaeo-lake (Lake Lisan) that filled much of the Jordan Valley during Late Pleistocene times. This paper describes that assemblage, comparing it to other Levantine Upper Palaeolithic assemblages of equivalent antiquity. The Mughr el-Hamamah Layer B assemblage appears most similar to Early Ahmarian assemblages, but it departs from typical such assemblages in ways that may reflect local conditions’ influence on human activities carried out in and near the cave. Mughr el-Hamamah raises new questions about changes in residential mobility, off-site provisioning and foraging activity, and on-site task diversity in the Early Upper Palaeolithic period.     

Sources of variation in carnivore tooth-mark frequencies in a modern spotted hyena (Crocuta crocuta) den assemblage,Amboseli Park,Kenya

Distributions and frequencies of carnivore tooth-marks on large mammal long-bone fragments are commonly used to infer the timing of hominin and carnivore access to prey resources in archaeofaunal assemblages. The strength of these inferences, however, is limited by a broad and currently inexplicable range of tooth-mark frequencies across experimental and archaeological assemblages. Controlling for this variation first requires that the sources be identified. Several sources of variation are examined here in an analysis of tooth-marked bone recovered from a modern spotted hyena (Crocuta crocuta) den assemblage in Amboseli Park, Kenya. Results indicate that tooth-mark frequencies: (1) depend on fragment size, (2) vary across mammals of different size classes, (3) are highly variable across equivalent portions of different long-bone elements, and (4) on certain long-bone portions are correlated with bone density and can be depressed in archaeological assemblages subjected to density-mediated attrition. Stronger inferences based on tooth-mark frequencies will require that such variation be taken into consideration, and methods for doing so are suggested.     

Big Game and Small Prey: Paleolithic and Epipaleolithic Economy from Valencia (Spain)

This study analyzes the faunal assemblages corresponding to the Middle and Upper Paleolithic and the Early Epipaleolithic for the central Spanish Mediterranean Region dated from between 135,000 and 9,000 B.P. To interpret these data, we employ a zooarchaeological and paleoeconomic perspective. Human large and small game hunting systems are analyzed through the identification of the origins of faunal assemblages, bone breackage patterns, and prey selection patterns. Our study shows that hunting systems changed significantly through time, which may be related to the characteristics of the Mediterranean ecosystem and, above all, to cultural changes.     

Archaeometry meets archaeozoology: Viking Haithabu and medieval Schleswig reconsidered

The two archaeofaunal assemblages from Haithabu (Viking period) and Schleswig (early Middle Age) belong to the largest such bone complexes ever recovered in Northern Europe. More than 800,000 animal bones were recovered from both sites in the course of the last three decades and investigated by traditional archaeozoological methods. More recently and independently from the latter, stable isotopic ratios of the light elements carbon, nitrogen, and oxygen were analyzed on a total of 440 animal bone finds representing 67 vertebrate species from both sites producing a data set comprising 1,760 individual isotopic ratios. In addition, stable strontium isotopic ratios were obtained from 68 animal teeth and bones of four species. In this paper, an attempt is made to synthesize archaeozoological and archaeometrical data on a larger scale. By becoming familiar with the specific potential of each method, we raise questions concerning not only palaeobiodiversity and palaeoecology of fish and birds in the area, trophic webs, and prey/predator relationships, but also human meat-provisioning, import of animals and animal products, and the reconstruction of hunting areas. This transdisciplinary approach led to an improvement in the quality of our interpretations and the validation of previously confirmed and/or rejected assumptions. It is still too early to expect answers to all these questions, but we are convinced that, in attempting to do so, the potential of coordinating such methods for future research will become evident, and we therefore strongly recommend an intensification of archaeometric analyses of archaeofaunal assemblages.     

Paleodietary analysis of a San Francisco Bay Area shellmound: stable carbon and nitrogen isotope analysis of late Holocene humans from the Ellis Landing site (CA-CCO-295)

The late Holocene archaeofaunal record of the San Francisco Bay Area demonstrates temporal declines in the abundance of low-cost, high-ranked marine and terrestrial resources. During later periods of occupation, faunal assemblages are often dominated by lower-ranked, higher-cost resources, suggesting an increase in diet breadth through time. Archaeological resource intensification models argue that this marks a late Holocene decline in foraging efficiency in the Bay Area, driven by human-induced harvest pressure. This study examines dietary change in the region using stable carbon and nitrogen isotope data from 65 human burials, spanning two temporal components (cal AD 55–890 and cal AD 762–1550) at the Ellis Landing site (CA-CCO-295). The strong linear relationship between collagen carbon and nitrogen isotope values reflects both marine and terrestrial food consumption, with individuals showing a high level of dietary variability at the site. No temporal trend or meaningful sex differences were found in isotope values. The widening of diet breadth predicted by the archaeofaunal record occurred prior to the occupation of the Ellis Landing shellmound site. The trend in exploiting a greater amount of terrestrial resources during the Middle and Late Period in central California was supported by isotopic evidence.     

Settlement Dimensions of the Vietnam-Born Population in Metropolitan Sydney

Australia has the largest Indochinese population per capita in the world, and Sydney supported sizeable populations from Vietnam in 1986. These populations became socially visible because of physical characteristics and clustering and have generated the response by some commentators that ghettos are forming. This paper examines the extent of residential concentration of Vietnamese-bom population in Sydney and changes over time in the distribution of settlement with reference to new immigration and internal migration. Gravitation group migration has reinforced concentration, but the result has not been ghettos, although there have been specific local problems of unemployment and housing access which may constrain residential mobility for some time. Manufacturing and residential restructuring have affected unemployment and have constrained dispersion. Institutional factors have reinforced concentration. Some residential mobility will be required for entry into a diversity of labour markets, if unemployment is to be reduced in the context of the restructuring of manufacturing production.     

Taphonomic analysis of rodent bone accumulations produced by Geoffroy's cat (Leopardus geoffroyi, Carnivora,Felidae) in Central Argentina

Rodent prey remains recovered from Geoffroy's cat (Leopardus geoffroyi) scats were analyzed in order to identify taphonomic features produced by this predator. The analysis includes a year-long sampling discriminated by the seasons. Modifications produced by digestion are heavy. Taphonomic variables in samples discriminated by the seasons did not show major differences with respect to the total sample; thus, scats collected in any season clearly show the modifications on the bones of prey rodents made by Geoffroy's cats. The results presented here and its interpretation could be extrapolated to an analysis of zooarchaeological or paleontological assemblages.     

Indirect Evidence of Falconry in Medieval Poland as Inferred from Published Zooarchaeological Studies

Although written sources indicate that falconry or hawking was very popular already in early medieval Poland, there is no hard zooarchaeological evidence that would support it. So far, only two studies based on animal remains suggested hawking in medieval Poland. The purpose of this study was to examine all published zooarchaeological reports from all archaeological sites in Poland and check for possible clues of hawking. Altogether, 281 remains of 12 species of diurnal birds of prey were recorded from 38 sites (49 time‐site units) in Poland beginning from the Middle Ages onward. The most frequently found was the goshawk Accipiter gentilis followed by the white‐tailed eagle Haliaeetus albicilla and the sparrowhawk Accipiter nisus. Clues of possible hawking include high status of sites, occurrence of species used in falconry outside their natural ranges, preponderance of females, the presence of sub‐adult specimens and others. It is suggested that surface damage observed on prey bones done by the hawks' beaks and claws may be an extra evidence of hawking. The present study strongly supports the notion that hawking was a very popular activity in medieval Poland. In the future, medieval bone assemblages should always be checked for clues of hawking. The present study may also be used as a source of raw data for other analyses because it includes detailed information on birds of prey from all zooarchaeological reports published so far in Poland. Copyright © 2015 John Wiley & Sons, Ltd.     

Zooarchaeology and Historical Archaeology: Progress and Prospects

This review assesses the development and current status of zooarchaeological research in historical archaeology. Analytical issues in recovery, identification, quantification and interpretation are discussed with particular reference to assemblages from historical sites. The results section summarizes the substantive contributions zooarchaeological studies of historical assemblages have made to our understanding of past diet, food production systems, social and cultural variation, and archaeological interpretations. The last four decades of research has provided a strong basis for future analyses that draw together diverse strands of zooarchaeological, historical, scientific, and anthropological evidence.     

Food for thought: using game cameras to better understand the movement of bones by scavenging in archaeological faunal assemblages

This paper questions our understanding of the movement of bones by animal scavengers in the archaeological record. Since assumptions regarding the effects of animal scavenging shape final interpretations of skeletal element frequencies in archaeological faunal assemblages, they are important for our understanding and reconstruction of ancient human behaviour. The results of a 4-year actualistic kangaroo scavenging study from Australia are used to question our understanding of the movement of the bone by contrasting visual data captured by motion-activated digital game cameras with traditional taphonomic studies using skeletal element frequencies. Game cameras are commonly used by ecologists to capture the behaviour of living species but have not yet been used in experimental archaeology where visually documenting animal scavenging behaviour can be used to understand the movement of carcasses and individual bones. Results suggest that traditional zooarchaeological analyses may not be accurate indicators of hunted versus scavenged prey in archaeological faunal assemblages. Moreover, they most certainly fail to document the entire suite of animals scavenging a carcass. These implications are discussed with particular reference to the ability to definitively ascertain the role of humans in the megafaunal extinction debate in Australia.     

Integrating Human-Animal Relationships into New Data on Aterian Complexity: a Paradigm Shift for the North African Middle Stone Age

In the last decade, research conducted in North Africa, particularly in northwestern Africa, has shed light on the key role that the region has played in improving our understanding of human evolution. Specifically, (1) the increased number of direct dates obtained with new methods has pushed back the age of the Aterian (~ 150,000 to 40,000 BP); (2) analyses of Aterian lithic assemblages have placed them within the range of variation of the Middle Stone Age; (3) analyses of associated human remains suggest that the makers of the Aterian are within the range of variation common among early modern human and present affinities with contemporary remains from the Levant (Qafzeh, Skhul); (4) zones of settlement, such as those in the present-day Sahara and coastal areas, and even the composition and demography of populations could have been influenced by specific climatic changes of the Late Pleistocene; and (5) the presence of blocks of pigment showing use-wear facets on their surfaces, the presence of pigments on artifacts, as well as osseous industry and earliest ornaments suggest complex behaviors among these populations. In this renewed approach to the Aterian, data from faunal analyses provide information on human-carnivore competition and the subsistence practices of hunter-gatherer groups. Taphonomic and zooarchaeological analyses suggest that humans were not the only large predators occupying caves, and that the hunter-gatherers of the Middle Stone Age exploited a wide range of environments, consuming terrestrial and coastal resources alike. Interdisciplinary confrontations highlight the apparent complexity of socioeconomic organization and the strategies of high levels of mobility that characterized Aterian groups.     

Subsistence variations and landscape use among maritime hunter-gatherers. A zooarchaeological analysis from the Beagle Channel (Tierra del Fuego,Argentina)

Changes in the exploitation of resources among prehistoric hunter-gatherer-fishers of the Beagle Channel (Southern South America) are examined in this paper. Archaeological investigations show the prevalence of maritime hunter-gatherer organization throughout the occupation of the region (ca. 6400 BP – 19th century). Notwithstanding, variations in the exploitation of different kind of animal resources have been detected, and a concomitant reorientation in the landscape use is then inferred. Zooarchaeological evidence from five shellmiddens is analyzed here: evenness measures and relative abundance prey types are used to evaluate such adjustments. Results indicate the major utilization of coastal and terrestrial ecozones and the predominance in the exploitation of mammals (pinnipeds and guanacos) in ancient occupations. While, during the Late Holocene an expansion in ranges of maritime mobility for subsistence is detected, which was associated with an increase in the representation of birds and fish in the zooarchaeological record.     

Palaeolithic Human Subsistence in Mount Carmel (Israel). A Taphonomic Assessment of Middle and Early Upper Palaeolithic Faunal Remains from Tabun,Skhul and el‐Wad

Detailed archaeozoological studies of Middle and Early Upper Palaeolithic (250–40 kya) faunal assemblages from Tabun, Skhul and el‐Wad caves, located in Mount Carmel (Israel), are presented here, expanding on the previous palaeontological analyses carried out by Garrod and Bate in the 1930´s and by Garrard five decades later. Despite the well‐known excavation bias of the older excavations, this study makes greater use of modern taphonomic techniques and statistical procedures, providing new palaeoeconomic insights into the subsistence behaviour of modern humans, Neanderthals and their predecessors in the Levant. The results show that anatomically modern humans (AMH) were more efficient at hunting and foraging in terms of human mobility as well as an incipient prey specialisation. In addition, different uses of the Tabun cave through time have been identified. Tabun B, initially considered as a palaeontological accumulation, was also occasionally accessed by Neanderthals in order to exploit naturally trapped ungulates, and there were sporadic and logistic occupations in Tabun D, one of the oldest Middle Palaeolithic assemblages in the area. Contrastingly, Tabun C showed a comparatively more residential use, which coincided with the new mobility capabilities of AMH. Whereas the intensity of human occupation at el‐Wad varied through time, levels G and F showed very brief occupations in comparison with levels D and E, which showed abundant evidence of human activity. Copyright © 2011 John Wiley & Sons, Ltd.     

Opportunistic versus target mode: Prey choice changes in central-western Korean prehistory

Human decision-making processes are usually hierarchical in that higher-level decisions impose constraints on lower-level decisions. As a result, prey choices during individual foraging trips are governed to a large degree by higher-level decisions regarding how to supply resources to satisfy demands, with higher-level decisions typically made prior to foraging trips. Resource selectivity and search bias sometimes take place in this context. By dividing resource procurement modes into opportunistic and target, I discuss how choice of mode on the basis of an overall economic plan affects prey choice during foraging trips and faunal assemblage composition resulting from those trips. An analysis of taxonomic diversity in shellmidden assemblages from the central-western Korean Late Chulmun Period (3500–1300 BC) and Middle and Late Mumun Period (700–100 BC) shows that Late Chulmun people adopted a target mode, while Middle/Late Mumun people adopted an opportunistic mode in their exploitation of marine resources. A decrease in the importance of marine resource in Middle/Late Mumun produced a change in taxonomic diversity by increasing the opportunistic cost of marine resource exploitation.     

Shifting stones and changing homes: using toolstone ratios to consider relative occupation span in the northwestern Great Basin

Like Paleoindian populations elsewhere in North America, Pre-Archaic groups in the Great Basin are assumed to have been highly mobile and far ranging. This view is commonly based on analyses of lithic technology and source provenance studies. While these approaches have added to our knowledge of Pre-Archaic lifeways, they have rarely focused on occupation span – an aspect of hunter–gatherer behavior directly related to mobility. Here, I use the proportions of local and nonlocal toolstone in Pre-Archaic and later Archaic assemblages to consider occupation span with the assumption that assemblages should become increasingly dominated by local materials as occupation span increases. The results suggest that residential mobility was high and occupations short before 7500 radiocarbon years ago. Conversely, between 7500 and 1300 radiocarbon years ago, residential mobility decreased and many locations were occupied for extended periods. Occupation span once again decreased as residential mobility increased after 1300 radiocarbon years ago. These trends were likely influenced by changes in the environmental and demographic climate of the Holocene.     

Pinniped behavior, foraging theory, and the depression of metapopulations and nondepression of a local population on the southern Northwest Coast of North America

Models derived from foraging theory suggest that high-value prey will be depressed (encounter rates will decrease) relative to low-value prey as human predation intensifies. Numerous case studies in ethnographic and zooarchaeological settings indicate depression of prey is very common. Exceptions to depression are few and have been cited as evidence of conservation and resource management. Reanalysis of pinniped and ungulate data from sites on the Oregon coast indicates that the metapopulation (entire population in the region) of otariids was depressed but a single local population of Steller sea lions exploited over a span of 600 years reached an equilibrium with human predation and was not subsequently depressed. This local population was not historically documented, and thus must have been extirpated by commercial sealing in the late nineteenth century. This singular case of prehistoric epiphenomenal conservation was an unintentional result of human predation that took advantage of the lack of escape behaviors among breeding male Steller sea lions.     

Explaining prehistoric variation in the abundance of large prey: A zooarchaeological analysis of deer and rabbit hunting along the Pecho Coast of Central California

Three main hypotheses are commonly employed to explain diachronic variation in the relative abundance of remains of large terrestrial herbivores: (1) large prey populations decline as a function of anthropogenic overexploitation; (2) large prey tends to increase as a result of increasing social payoffs; and (3) proportions of large terrestrial prey are dependent on stochastic fluctuations in climate. This paper tests predictions derived from these three hypotheses through a zooarchaeological analysis of eleven temporal components from three sites on central California’s Pecho Coast. Specifically, we examine the trade-offs between hunting rabbits (Sylvilagus spp.) and deer (Odocoileus hemionus) using models derived from human behavioral ecology. The results show that foragers exploited a robust population of deer throughout most of the Holocene, only doing otherwise during periods associated with climatic trends unfavorable to larger herbivores. The most recent component (Late Prehistoric/Contact era) shows modest evidence of localized resource depression and perhaps greater social benefits from hunting larger prey; we suggest that these final changes resulted from the introduction of bow and arrow technology. Overall, results suggest that along central California’s Pecho Coast, density independent factors described as climatically-mediated prey choice best predict changes in the relative abundance of large terrestrial herbivores through the Holocene.     

Taphonomy and zooarchaeology of the Upper Palaeolithic cave of Dzudzuana,Republic of Georgia

We present the results of a detailed taphonomic and zooarchaeological study of the faunal remains from the Upper Palaeolithic layers of Dzudzuana Cave, Republic of Georgia. This study presents the first carefully analysed Upper Palaeolithic faunal assemblage from the southern Caucasus and thus serves as a significant point of reference for inter‐regional studies of Upper Palaeolithic subsistence in Eurasia. A series of intra‐site taphonomic comparisons are employed to reconstruct the depositional history of the bone assemblages within the different occupational phases at the site and to investigate subsistence, meat procurement and bone‐processing strategies. Caucasian tur (Capra caucasica), aurochs (Bos primigenius) and steppe bison (Bison priscus) were the major prey species throughout the Upper Palaeolithic. Their frequencies do not change significantly over time, and nor does bone preservation vary by layer. The assemblage is characterised by significant density‐mediated biases, caused by both human bone‐processing behaviours and in situ post‐burial bone attrition. Bone marrow extraction produced large numbers of unidentified bone fragments, many exhibiting green bone fractures. The density and size of bone assemblages and the extent of fragmentation indicate that Dzudzuana Cave was repeatedly occupied by Upper Palaeolithic foragers over many years. Skeletal part representation and butchery marks from all stages of carcass processing suggest that prey occasionally underwent field butchery. Intra‐site taphonomic comparisons highlight uniform patterns of cultural and economic behaviours related to food procurement and processing strategies. Copyright © 2007 John Wiley & Sons, Ltd.     

Faunal exploitation and agricultural transitions in the South American agricultural limit

The southern Mendoza Province of Argentina constitutes the southernmost limit of Andean agriculture in South America. Archaeofaunal assemblages from sites in this region show a pattern of changing taxonomic diversity concomitant with the first appearance of domesticated plants. This pattern is characterised by an increase in prey diversity before the appearance of the first cultigens and a decrease in diversity after their arrival. Furthermore, the animal taxa exploited when domesticated plants entered the diet indicate a focus on big game. The pattern observed in southern Mendoza is in accord with ethnographic and archaeological models generated by faunal research worldwide. Copyright © 2008 John Wiley & Sons, Ltd.